Cristate and plumose ferns are another instance. Half races or rare
accidental cleavages seem to be as common with ferns as cultivated
double races, which are very rich in beautiful crests. But much depends
on cultivation. It seems that the spores of crested leaves are more apt
to reproduce the variety than those of normal leaves, or even of normal
parts of the same leaves. But the experiments on which this assertion is
made are old and should be repeated. Other cases of cleft leaves should
also be tested. Ascidia are far more common than is usually believed.
Rare instances point [428] to poor races, but the magnolias and
lime-trees are often so productive of ascidia as to suggest the idea of
ever-sporting varieties. I have seen many hundred ascidia on one
lime-tree, and far above a hundred on the magnolia. They differ widely
in size and shape, including in some cases two leaves instead of one, or
are composed of only half a leaf or of even still a smaller part of the
summit. Rich ascidia-bearing varieties seem to offer notable
opportunities for scientific pedigree-cultures.
Union of the neighboring fruits and flowers on flower-heads, of the rays
of the umbellifers or of the successive flowers of the racemes of
cabbages and allied genera, seem to be rare. The same holds good for the
adhesion of foliar to axial organs, of branches to stems and other cases
of union. Many of these cases return regularly in each generation, or
may at least be seen from time to time in the same strains.
Proliferation of the inflorescence is very common and changes in the
position of staminate and pistillate flowers are not rare. We find
starting points for new investigations in almost any teratological
structure. Half-races and double-races are to be distinguished and
isolated in all cases, and their hereditary qualities, the periodicity
of the recurrence of the anomaly, the dependency on external
circumstances [429] and many other questions have to be answered.
Here is a wide field for garden experiments easily made, which might
ultimately yield much valuable information on many questions of heredity
of universal interest.
[430]
LECTURE XV
DOUBLE ADAPTATIONS
The chief object of all experimentation is to obtain explanations of
natural phenomena. Experiments are a repetition of things occurring in
nature with the conditions so guarded and so closely followed that it is
possible to make a clear analysis of facts and their causes, it being
rightfully assumed that the laws are the same in both cases.
Experiments on heredity and the experience of the breeder find their
analogy in the succession of generations in the wild state. The
stability of elementary species and of retrograde varieties is quite the
same under both conditions. Progression and retrogression are narrowly
linked everywhere, and the same laws govern the abundance of forms in
cultivated and in wild plants.
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