Such varieties however, do not exist at present. The ordinary twisted
races on the other hand, are found in the wild state and have only to be
isolated and cultivated to yield large numbers [447] of twisted
individuals. In nature they are able to maintain themselves during long
centuries, quite as well as normal species and varieties. But they owe
this quality entirely to their dimorphous character. A twisted race of
teasels might consist of successive generations of tall atavistic
individuals, and produce yearly some twisted specimens, which might be
destroyed every time before ripening their seeds. Reasoning from the
evidence available, and from analogous cases, the variety would, even
under such extreme circumstances, be able to last as long as any other
good variety or elementary species. And it seems to me that this
explanation makes clear how it is possible that varieties, which are
potentially rich in their peculiar monstrosity, are discovered from time
to time among plants when tested by experimental methods.
Granting these conclusions, monstrosities on the one side, and
dimorphous wild species on the other, constitute the most striking
examples of the inheritance of latent characters.
The bearing of the phenomena of dimorphism upon the principles of
evolution formulated by Lamarck, and modified by his followers to
constitute Neo-Lamarckianism, remains to be considered. Lamarck assumed
that the external conditions directly affected the organisms in [448]
such a way as to make them better adapted to life, under prevailing
circumstances. Nageli gave to this conception the name "Theory of direct
causation" (Theorie der directen Bewirkung), and it has received the
approval of Von Wettstein, Strasburger and other German investigators.
According to this conception a plant, when migrating from lowlands into
the mountains would slowly be changed and gradually assume alpine
habits. Once acquired this habit would become fixed and attain the rank
of specific characters. In testing this theory by field-observations and
culture-experiments, the defenders of the Nagelian principle could
easily produce evidence upon the first point. The change of
lowland-plants into alpine varieties can be brought about in numerous
cases, and corresponding changes under the influence of soil, or
climate, or life-conditions are on record for the most various
characters and qualities.
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