The first record of the peloric toad-flax is that of Zioberg, a student
of Linnaeus, who found it in the neighborhood of Upsala. This curious
discovery was described by Rudberg in his dissertation in the year 1744.
Soon afterwards other localities were discovered by Link near Gottingen
in Germany about 1791 and afterwards [467] in the vicinity of Berlin, as
stated by Ratzeburg, 1825. Many other localities have since been
indicated for it in Europe, and in my own country some have been noted
of late, as for instance near Zandvoort in 1874 and near Oldenzaal in
1896. In both these last named cases the peloric form arose
spontaneously in places which had often been visited by botanists before
the recorded appearance, and therefore, without any doubt, they must
have been produced directly and independently by the ordinary species
which grows in the locality. The same holds good for other occurrences
of it. In many instances the variety has been recorded to disappear
after a certain lapse of time, the original specimens dying out and no
new ones being produced. _Linaria_ is a perennial herb, multiplying
itself easily by buds growing on the roots, but even with this means of
propagation its duration seems to have definite limits.
There is one other important point arguing strongly for the independent
appearance of the peloric form in its several localities. It is the
difficulty of fertilization and the high degree of sterility, even if
artificially pollinated. Bees and bumble-bees are unable to crawl into
the narrow tubular flowers, and to bring the fertilizing pollen to the
stigma. Ripe capsules with seeds [468] have never been seen in the wild
state. The only writer who succeeded in sowing seeds of the peloric
variety was Wildenow and he got only very few seedlings. But even in
artificial pollination the result is the same, the anthers seeming to be
seriously affected by the change. I tried both self-fertilization and
cross-pollination, and only with utmost care did I succeed in saving
barely a hundred seeds. In order to obtain them I was compelled to
operate on more than a thousand flowers on about a dozen peloric plants.
The variety being wholly barren in nature, the assumption that the
plants in the different recorded localities might have a common origin
is at once excluded. There must have been at least nearly as many
mutations as localities. This strengthens the hope of seeing such a
mutation happen in one's own garden. It should also be remembered that
peloric flowers are known to have originated in quite a number of
different species of _Linaria_, and also with many of the allied species
within the range of the Labiatiflorae.
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