Thirdly, the question arises, whether the mutation is complete, not only as to the morphologic character, but also as to the hereditary constitution of the mutated individuals. But here unfortunately the high degree of sterility of the peloric plants, as previously noted, makes the experimental evidence a thing of great difficulty. During the course of several years I isolated and planted together the peloric individuals already mentioned, all in all some twenty plants. Each individual was nearly absolutely sterile when treated with its own pollen, and the aid of insects was of no avail. I intercrossed my plants artificially, and pollinated more than a thousand flowers. Not a single one gave a normal fruit, but some small and nearly rudimentary capsules were produced, bearing a few seeds. From these I had 119 flowering plants, out of which 106 were peloric and 13 one-spurred. The great majority, some 90%, were thus shown to be true to their new type. Whether the 10% reverting ones were truly atavists, or whether they were [478] only vicinists, caused by stray pollen grains from another culture, cannot of course be decided with sufficient certitude. Here I might refer to the observations concerning the invisible dimorphous state of the flowers of the normal toad-flax. Individuals of the same type, when fertilized with each other, are nearly, but not absolutely, sterile. The yield of seeds of my peloric plants agrees fairly well with the harvest which I have obtained from some of the nearly sterile pairs of individuals in my former trial. Hence the suggestion is forced upon us that perhaps, owing to some unknown cause, all the peloric individuals of my experiment belonged to one and the same type, and were sterile for this reason only. If this is true, then it is to be presumed that all previous investigators have met the same condition, each having at hand only one of the two required types. And this discussion has the further advantage of showing the way, in which perhaps a full and constant race of peloric toad-flaxes may be obtained. Two individuals of different type are required to start from. They seem as yet never to have arisen from one group of mutations. But if it were possible to combine the products of two mutations obtained in different countries and under different conditions, there would be a chance [479] that they might belong to the supposed opposite types, and thus be fertile with one another. My peloric plants are still available, and the occurrence of this form elsewhere would give material for a successful experiment. The probability thereof is enhanced by the experience that my peloric plants bear large capsules and a rich harvest of seeds when fertilized from plants of the normal one-spurred race, while they remain nearly wholly barren by artificial fertilization with others. I suppose that they are infertile with the normal toad-flaxes of their own sexual disposition, but fertile with those of the opposite constitution.