This last is therefore, of necessity, always continued by means of
individuals whose deviation from the mean average is the least. But in
many cases the varieties are so highly differentiated that selection has
become quite superfluous for practical purposes. I have already
discussed the question as to the actual moment, in which the change of
the _grandiflorum_ variety into the new _plenum_ form must be assumed to
have taken place. In this respect some stress is to be laid on the fact
that the improvement through selection has been gradual and continuous,
though very rapid from the first moment. But with the appearance of the
first stray rays within the disk, this continuity suddenly changed. All
the children of this original mutated plant showed the new character,
the rays within the disk, without exception. Not on all the heads, nor
even on the majority of the heads on some individuals, but on some heads
all gave clear proof of the possession of the new attribute. This was
present in all the representatives of the new race, and had never been
seen in any of their parents and grandparents. Here there was evidently
a sudden leap, at least in the external form of the plants. And it seems
to me to be the most simple conception, [505] that this visible leap
directly corresponded to that inner change, which brought about the
complete inheritability of the new peculiarity. It is very interesting
to observe how completely my experience agrees with the results of the
observations of breeders at large. No doubt a comparison is difficult,
and the circumstances are not adequate to a close study.
Isolation and selection have been applied commonly only so far as was
consistent with the requirements of practical horticulture, and of
course a determination of the hereditary percentage was never made. The
disregard of this feature made necessary a greater length of time and a
larger number of generations to bring about the desired changes.
Notwithstanding this, however, it has been seen that double varieties
are produced suddenly. This may have occurred unexpectedly or after a
few years' effort toward the end desired. Whether this sudden appearance
is the consequence of a single internal differentiating step, or of the
rapid succession of lesser changes, cannot yet be made out. The extreme
variability of double flowers and the chance of their appearance with
only slight indications of the previous petaloid alterations of a few
stamens may often result in their origin being overlooked, while
subsequent generations may come in for full notice. [506] In the greater
number of cases recorded it remains doubtful whether the work said to be
done to obtain a new double variety was done before the appearance of
these preliminary indications or afterward.
In the first case, it would correspond with our selection of large
numbers of florets in the outer rays, in the second however, with the
ordinary purification of new races from hybrid mixtures.
Public-domain text, read in full here on John Shaqi.
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