It is readily granted that the constant condition of species is the
normal one, and that mutating periods must be the exception. This fact
does not tend to increase our prospect of discovering a species in a
state of mutability. Many species will have to be tested before finding
an instance. On the other hand, a direct trial seems to be the only way
to reach the goal. No such special guides as those that led us to the
choice of pelories and double flowers are available. The only indication
of value is the presumption that a condition of mutability might be
combined with a general state of variability at large, and that groups
of plants of very uniform features might be supposed to be constant in
this respect too. On the contrary, anomalies and deviations if existent
in the members of one strain, or found together in one native locality
of a species, might be considered as an indication in the desired
direction.
Few plants vary in the wild state in such a [520] measure as to give
distinct indications. All have to be given a trial in the garden under
conditions as similar as possible to their natural environments.
Cultivated plants are of course to be excluded. Practically they have
already undergone the experience in question and can not be expected to
change their habits soon enough. Moreover they are often of hybrid
origin. The best way is to experiment with the native plants of one's
own country.
I have made such experiments with some hundred species that grow wild in
Holland. Some were very variable, as for instance, the jointed charlock
(_Raphanus Raphanistrum_) and the narrow-leaved plantain (_Plantago
lanceolata_). Others seemed more uniform, but many species, collected
without showing any malformation, subsequently produced them in my
garden, either on the introduced plants themselves or among their
offspring. From this initial material I have procured a long series of
hereditary races, each with some peculiar anomaly for its special
character. But this result was only a secondary gain, a meager
consolation for the negative fact that no real mutability could be
discovered.
My plants were mostly annuals or biennials, or such perennials as under
adequate treatment might produce flowers and seeds during their [521]
first summer. It would be of no special use to enumerate them. The
negative result does not apply to the species as such, but only to the
individual strain, which I collected and cultivated. Many species, which
are quite constant with us, may be expected to be mutable in other parts
of their range.
Only one of all my tests met my expectations. This species proved to be
in a state of mutation, producing new elementary forms continually, and
it soon became the chief member of my experimental garden. It was one of
the evening primroses.
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