Secondly the instability seems to be a constant quality, although the
words themselves are at first sight, contradictory. I mean to convey the
conception that the degree of instability remains unchanged during
successive generations. This is a very curious fact, and strongly
reminds us of the hereditary conditions of striped-flower varieties.
But, on the contrary, the atavists, which are here the individuals with
the stature and the characteristics of the _lamarckiana_, have become
_lamarckiana_s in their hereditary qualities, too. If their seed is
saved and sown, their progeny does not contain any _scintillans_, or at
least no more than might arise by ordinary mutations.
One other inconstant new species is to be noted, but as it was very rare
both in the field and in my cultures, and as it was difficult of
cultivation, little can as yet be said about it. It is the _Oenothera
elliptica_, with narrow elliptical leaves and also with elliptical
petals. It repeats [546] its type only in a very small proportion of its
seed.
All in all we thus have a group of a dozen new types, springing from an
original form in one restricted locality, and seen to grow there, or
arising in the garden from seeds collected from the original locality.
Without any doubt the germs of the new types are fully developed within
the seed, ready to be evolved at the time of germination. More favorable
conditions in the field would no doubt allow all of the described new
species to unfold their attributes there, and to come into competition
with each other and with the common parents. But obviously this is only
of secondary importance, and has no influence on the fact that a number
of new types, analogous to the older swarms of _Draba_, _Viola_ and of
many other polymorphous species, have been seen to arise directly in the
wild state.
[547]
LECTURE XIX
EXPERIMENTAL PEDIGREE-CULTURES
The observation of the production of mutants in the field at Hilversum,
and the subsequent cultivation of the new types in the garden at
Amsterdam, gives ample proof of the mutability of plants. Furthermore it
furnishes an analogy with the hypothetical origin of the swarms of
species of _Draba_ and _Viola_. Last but not least important it affords
material for a complete systematic and morphologic study of the newly
arisen group of forms.
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