_Albida_, _nanella_ and _rubrinervis_ appeared in large numbers, and
even _scintillans_, of which I had but a single plant in the previous
generation, was repeated sixfold.
New forms did not arise, and the capacity of my strain seemed exhausted.
This conclusion was strengthened by the results of the next three
generations, which were made on a much smaller scale and yielded the
same, or at least the mutants most commonly seen in previous years.
Instead of giving the figures for these last two years separately, I
will now summarize my whole experiment in the form of a pedigree. In
this the normal _lamarckiana_ was the main line, and seeds were only
sown from plants after sufficient isolation either of the plants
themselves, or in the latter years by means of paper bags enclosing the
inflorescences. I have given the number of seedlings of _lamarckiana_
which were examined each year in the table below. Of course by far the
largest number of them were [556] thrown away as soon as they showed
their differentiating characters in order to make room for the remaining
ones. At last only a few plants were left to blossom in order to
perpetuate the race. I have indicated for each generation the number of
mutants of each of the observed forms, placing them in vertical columns
underneath their respective heads. The three first generations were
biennial, but the five last annual.
PEDIGREE OF A MUTATING FAMILY
OF _OENOTHERA LAMARCKIANA_ IN THE
EXPERIMENTAL GARDEN AT AMSTERDAM
Gener: O.gig. albida obl. rubrin. Lam. nanella lata. scint.
VIII. 5 1 0 1700 21 1
VII. 9 0 3000 11
VI. 11 29 3 1800 9 5 1
V. 25 135 20 8000 49 142 6
IV. 1 15 176 8 14000 60 73 1
III. 1 10000 3 3
II. 15000 5 5
I. 9
It is most striking that the various mutations of the evening-primrose
display a great degree of regularity. There is no chaos of forms, no
indefinite varying in all degrees and in all directions. Quite on the
contrary, it is at once evident that very simple rules govern the whole
phenomenon.
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