On the other hand the mutants, that constitute the first representatives
of their race, exhibit all the attributes of the new type in full
display at once. No series of generations, no selection, [559] no
struggle for existence are needed to reach this end. In previous
lectures I have mentioned that I have saved the seeds of the mutants
whenever possible, and have always obtained repetitions of the prototype
only. Reversions are as absolutely lacking as is also a further
development of the new type. Even in the case of the inconstant forms,
where part of the progeny yearly return to the stature of _lamarckiana_,
intermediates are not found. So it is also with _lata_, which is
pistillate and can only be propagated by cross-fertilization. But though
the current belief would expect intermediates at least in this case,
they do not occur. I made a pedigree-culture of lata during eight
successive generations, pollinating them in different ways, and always
obtained cultures which were partly constituted of _lata_ and partly of
_lamarckiana_ specimens. But the _lata_s remained _lata_ in all the
various and most noticeable characters, never showing any tendency to
gradually revert into the original form.
Intermediate forms, if not occurring in the direct line from one species
to another, might be expected to appear perhaps on lateral branches. In
this case the mutants of one type, appearing in the same year, would not
be a pure type, but would exhibit different degrees of deviation from
the parent. The best would then have to [560] be chosen in order to get
the new type in its pure condition. Nothing of the kind, however, was
observed. All the _oblonga_-mutants were pure _oblongas_. The pedigree
shows hundreds of them in the succeeding years, but no difference was
seen and no material for selection was afforded. All were as nearly
equal as the individuals of old elementary species.
II. New forms spring laterally from the main stem.
The current conception concerning the origin of species assumes that
species are slowly converted into others. The conversion is assumed to
affect all the individuals in the same direction and in the same degree.
The whole group changes its character, acquiring new attributes. By
inter-crossing they maintain a common line of progress, one individual
never being able to proceed much ahead of the others.
Public-domain text, read in full here on John Shaqi.
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