This is a very curious fact. It embraces two minor points, viz: the
multitude of similar mutants in the same year, and the repetition
thereof in succeeding generations. Obviously there must be some common
cause. This cause must be assumed to lie dormant in the _Lamarckiana_s
of my strain, and probably in all of them, as no single parent-plant
proved ever to be wholly destitute of mutability. Furthermore the
different causes for the sundry mutations must lie latent together in
the same parent-plant. They obey the same general laws, become active
under similar conditions, some of them being more easily awakened than
others. The germs of the _oblonga_, _lata_ and _nanella_ are especially
irritable, and are ready to spring into activity at the least summons,
while those of _gigas_, _rubrinervis_ and _scintillans_ are far more
difficult to arouse.
These germs must be assumed to lie dormant during many successive
generations. This is especially evident in the case of _lata_ and
_nanella__, which appeared in the first year of the pedigree culture and
which since have been repeated yearly, and have been seen to arise by
mutation [567] also during the last season (1903). Only _gigas_ appeared
but once, but then there is every reason to assume that in larger
sowings or by a prolongation of the experiments it might have made a
second appearance.
Is the number of such germs to be supposed to be limited or unlimited?
My experiment has produced about a dozen new forms. Without doubt I
could easily have succeeded in getting more, if I had had any definite
reason to search for them. But such figures are far from favoring the
assumption of indefinite mutability. The group of possible new forms is
no doubt sharply circumscribed. Partly so by the morphologic
peculiarities of _lamarckiana_, which seem to exclude red flowers,
composite leaves, etc. No doubt there are more direct reasons for these
limits, some changes having taken place initially and others later,
while the present mutations are only repetitions of previous ones, and
do not contribute new lines of development to those already existing.
This leads us to the supposition of some common original cause, which
produced a number of changes, but which itself is no longer at work, but
has left the affected qualities, and only these, in the state of
mutability.
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