_Epilobium hirsutum cruciatum_ was discovered by John Rasor near
Woolpit, Bury St. Edmunds, in England. It flowered in one spot,
producing about a dozen stems, among large quantities of the
parent-species, which is very common there, as it is elsewhere in
Europe. This species is a perennial, multiplying itself by underground
runners, and the stems of the new variety were observed to stand so
close to each other that they might be considered as the shoots of one
individual. In this case this specimen might probably be the original
mutant, as the variety had not been seen on that spot in previous years,
even as it has not been found elsewhere in the vicinity.
Intermediates were not observed, though the difference is a very
striking one. In the cruciate flowers the broad and bright purple petals
seem at first sight to be wholly wanting. They are too weak to expand
and to reflex the calyx [589] as in the normal flowers of the species.
The sepals adhere to one another, and are only opened at their summit by
the protruding pistils. Even the stamens hardly come to light. At the
period of full bloom the flowers convey only the idea of closed buds
crowned by the conspicuous white cross of the stigma. Any intermediate
form would have at once betrayed itself by larger colored petals, coming
out of the calyx-sheath. The cruciate petals are small and linear and
greenish, recalling thereby the color of the sepals.
Mr. Rasor having sent me some flowers and some ripe capsules of his
novelty, I sowed the latter in my experimental garden, where the plant
flowered in large numbers and with many thousands of flowers both in
1902 and 1903. All of these plants and all of these flowers repeated the
cruciate type exactly, and not the slightest impurity or tendency to
partial reversion has been observed.
Thus true and constant cruciate varieties have been produced from
accidentally observed initial plants, and because of their very curious
characters they will no doubt be kept in botanical gardens, even if they
should eventually become lost in their native localities.
At this point I might note another observation made on the wild species
of _Oenothera cruciata_ [590] from the Adirondacks. Through the kindness
of Dr. MacDougal, of the New York Botanical Garden, I received seeds
from Sandy Hill near Lake George. When the plants, grown from these
seeds, flowered, they were not a uniform lot, but exhibited two distinct
types. Some had linear petals and thin flower-buds, and in others the
petals were a little broader and the buds more swollen. The difference
was small, but constant on all the flowers, each single plant clearly
belonging to one or the other of the two types. Probably two elementary
species were intermixed here, but whether one is the systematic type and
the other a mutation, remains to be seen.
Public-domain text, read in full here on John Shaqi.
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