Weeping varieties afford similar instances. _Sophora japonica pendula_
originated about 1850, and _Gleditschia triacanthos pendula_ some time
later in a nursery at Chateau-Thierry (Aisne, France). In the year 1821
the bird's cherry, or _Prunus Padus_, produced a weeping variety, and in
1847 the same mutation was observed for the allied _Prunus Mahaleb_.
Numerous other instances of the sudden origin of weeping trees, both of
conifers and of others, have been brought together in Korshinsky's
paper. This striking type of variation includes perhaps the best
examples of the whole historical evidence. As a rule they appear in
large sowings, only one, or only a few at a time. Many of them have not
been observed during their youth, but only after having been planted out
in parks and forests, since the weeping characters show only after
several years.
The monophyllous bastard-acacia originated in the same way. Its
peculiarities will be dealt with on another occasion, but the
circumstances of its birth may as well be given here. In 1855 in the
nursery of Deniau, at Brain-sur-l'Authion (Maine et Loire), it appeared
in a lot of [618] seedlings of the typical species in a single
individual. This was transplanted into the Jardin des Plantes at Paris,
where it flowered and bore seeds in 1865. It must have been partly
pollinated by the surrounding normal representatives of the species,
since the seeds yielded only one-fourth of true offspring. This
proportion, however, has varied in succeeding years. Briot remarks that
the monophyllous bastard acacia is liable to petaloid alterations of its
stamens, which deficiency may encroach upon its fertility and
accordingly upon the purity of its offspring.
Broom-like varieties often occur among trees, and some are known for
their very striking reversions by buds, as we have seen on a previous
occasion. They are ordinarily called pyramidal or fastigiate forms, and
as far as their history goes, they arise suddenly in large sowings of
the normal species. The fastigiate birch was produced in this way by
Baumann, the _Abies concolor fastigiata_ by Thibault and Keteleer at
Paris, the pyramidal cedar by Paillat, the analogous form of
_Wellingtonia_ by Otin. Other instances could easily be added, though of
course some of the most highly prized broom-like trees are so old that
nothing is known about their origin. This, for instance, is the case
with the pyramidal yew-tree, _Taxus baccata fastigiata_. [619] Others
have been found wild, as already mentioned in a former lecture.
An analogous case is afforded by the purpleleaved plums, of which the
most known form is Prunus Pissardi. It is said to be a purple variety of
_Prunus cerasifera_, and was introduced at the close of the seventies
from Persia, where it is said to have been found in Tabris. A similar
variety arose independently and unexpectedly in the nursery of Spath,
near Berlin, about 1880, but it seems to differ in some minor points
from the Persian prototype.
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