The Italian poplar affords another instance. It is considered by some
authors as a distinct species, _Populus italica_, and by others as a
[624] broom-like variety of the _Populus nigra_, from which it is
distinguished by its erect branches and other characters of minor
importance. It is often called the pyramidal or fastigiate poplar. Its
origin is absolutely unknown and it occurs only in the cultivated state.
In Italy it seems to have been cultivated from the earliest historical
times, but it was not introduced into other countries till the
eighteenth century. In 1749 it was brought into France, and in 1758 into
England, and to day it may be seen along roads throughout central Europe
and in a large part of Asia. But the most curious fact is that it is
only observed in staminate specimens; pistillate trees have not been
found, although often sought for. This circumstance makes it very
probable that the origin of the broom-like poplar was a sudden mutation,
producing only one individual. This being staminate, it has been
propagated exclusively by cuttings. It is to be admitted, however, that
no material evidence is at hand to prove that it is not an original wild
species, the pistillate form of which has been lost by vegetative
multiplication. One form only of many dioecious plants is to be found in
cultivation, as, for instance some South American species of _Ribes_.
Total lack of historical evidence concerning [625] the origin of a
variety has sometimes been considered as sufficient proof of a sudden
origin. The best known instance is that of the renowned cactus-dahlia
with its recurved instead of incurved ray-florets. It was introduced
from Mexico into the Netherlands by Van den Berg of Jutphaas, under the
following remarkable circumstances. In the autumn of 1872 one of his
friends had sent him a small case, containing seeds, bulbs and roots
from Mexico. From one of these roots a _Dahlia_ shoot developed. It was
cultivated with great care and bloomed next year. It surprised all who
saw it by the unexpected peculiarity of its large rich crimson flowers,
the rays of which were reversed tubular. The margins of the narrow rays
were curved backwards, showing the bright color of the upper surface. It
was a very showy novelty, rapidly multiplied by cuttings, and was soon
introduced into commerce. It has since been crossed with nearly all
other available varieties of the _Dahlia_, giving a large and rich group
of forms, bound together by the curious curling of the petals. It has
never been observed to grow in Mexico, either wild or in gardens, and
thus the introduced individual has come to be considered as the first of
its race.
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