The steady cooperation of progression and retrogression is one of the
important principles of organic evolution. I have dwelt upon this point
more than once in previous lectures. I have tried to show that both in
the more important lines of the general pedigree of the vegetable
kingdom, and in the numerous lateral branches ending in the genera and
species within the families, progression and retrogression are nearly
always at work together. Your attention has been directed to the
monocotyledons as an example, where retrogression is everywhere so
active that it can almost be said to be the prevailing movement.
Reduction in the vegetative and generative organs, in the anatomical
structure and growth of the stems, and in sundry other ways is the
method by which the monocotyledons have originated as a group from their
supposed ancestors among the lower dicotyledonous families.
Retrogression is the leading idea in the larger families of the group,
[631] as for instance in the aroids and the grasses. Retrograde
evolution is also typical in the highest and most highly differentiated
family of the monocotyledons, the orchids, which have but one or two
stamens. In the second place I have had occasion more than once to
assert that retrogression, though seemingly consisting in the
disappearance of some quality, need not, as a rule, be considered as a
complete loss. Quite on the contrary, it is very probable that real
losses are extremely rare, if not wholly lacking. Ordinarily the loss is
only apparent, the capacity becomes inactive only, but is not destroyed.
The character has become latent, as it is commonly stated, and therefore
may return to activity and to the full display of its peculiarity,
whenever occasion offers.
Such a return to activity was formerly called atavism. But as we have
seen, when dealing with the phenomena of latency at large, sundry cases
of latency are to be distinguished, in order to get a clear insight into
these difficult processes.
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