Intermediate forms with respect to stature may also be seen. The
opium-poppy, the snapdragon, peas, the _Nicandra_, and many other
garden-plants have not only dwarf varieties, but also some of
intermediate height. These, though they are intermediate between the
tall and dwarf types, cannot be considered as transitions, as between
them and the extremes, intermediates are, as a rule wholly lacking.
Instances of the same occurrence of three types may be seen in the seeds
of maize ("Cuzco," "Horse-dent" and "Gracillima") of beans and some
other plants. The _Xanthium Wootoni_, above referred to, with only part
of the prickles of Xanthium commune is also a very curious instance of
the demonstration of the compound nature of a character.
Summarizing the conclusions that may be drawn from the evidence given in
this lecture, we have seen that varieties differ from elementary species
in that they do not possess anything really new. They originate for the
greater part in a negative way, by the apparent loss of some quality,
and rarely in a positive manner by acquiring a character, already seen
in allied species. These characters are not of the nature of [153]
morphologic entities, but are to be considered as physiologic units,
present in all parts of the organisms, and manifesting themselves where
ever occasion is afforded. They are units in the sense that they may
appear and disappear singly. But very often they are combined to yield
compound characters, which are capable of analysis. Opportunities for
such an analysis are afforded by these groups of cultivated varieties,
of which some members show a single distinguishing quality, or a number
of them.
[154]
LECTURE VI
STABILITY AND REAL ATAVISM
It is generally believed that varieties are principally distinguished
from species by their inconstancy. This conception is derived from some
special cases and transferred to others, and in its common form this
belief must have originated from the confusion which exists as to the
meaning of the term variety. It is true that vegetative varieties as a
rule run back, when propagated by seeds; they are an obvious instance of
inconstancy. In the second place we have considered the group of
inconstant or sporting varieties, which of course we must exclude when
studying the stability of other types. However, even these sporting
varieties are unstable only to a certain degree, and in a broader sense
will prove to be as true to their character as the most constant types.
Having separated these two groups, which include also the wide range of
hybrid forms, we may next consider only those varieties of pure origin,
and ordinarily propagated by seeds, [155] which have been discussed in
former chapters. Their general character lies in their fidelity to type,
and in the fact that this is single, and not double, as in the sporting
varieties.
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