The best proof is afforded by the varieties which grow wild in
localities where they are quite isolated from the species, and where for
this reason, no possibility of crossing disturbs the significance of the
proof. As one instance the rayless form of the wild camomile, or the
_Matricaria Chamomilla discoidea_ may be mentioned. Many systematists
have been so strongly [157] impressed with its absolute constancy and
its behavior as an ordinary species, that they have elevated it, as it
is called, to the rank of a species. As such it is described under the
name of _Matricaria discoidea_ DC. It is remarkable for its rapid and
widespread distribution, as of late years it has become naturalized in
different parts of America and of Europe, where it is to be seen
especially in France and in Norway. Experimentally I raised in
succeeding years between 1000 and 2000 seedlings, but observed no trace
of reversion, either in the strongest or in the numerous very small and
weak individuals which appeared in the cultures.
The tansy-ragwort or _Senecio Jacobaea_ may be chosen as a second
instance. It is a perennial herb with short rootstocks and stout stems
bearing numerous short-peduncled heads in large compact corymb; it
multiplies itself abundantly by seeds and is very common on the sand
dunes of Holland. It has two forms, differing only in the occurrence or
the lack of the ray florets. But these two varieties occupy different
localities and are even limited to different provinces. As far as I have
been able to ascertain on numerous excursions during a series of years,
they never sport, and are only intermingled on the outskirts of their
habitats. The rayless form is generally considered as the [158] variety
but it is quite as stable as the radiate species.
The radiate varieties of marigold, quoted in a former lecture, seem to
be equally constant, when growing far away from their prototypes. I
sowed the seeds of a single plant of the radiate form of _Bidens
cernua_, and found all of the seedlings came true, and in the next year
I had from their seed between 2,000 and 3,000 flowering individuals, all
equally radiate. Many species of composites have been tried, and they
are all constant. On the other hand rare sports of this kind have been
observed by Murr and other authors.
Many kinds of vegetables and of fruits give instances of stability.
White strawberries, green grapes, white currants, crisped lettuce,
crisped parsley and some other crisped forms may be cited. The spinage
without prickles is a widely known instance. White-flowered flax never
reverts to the blue prototype, if kept pure. Sugar-peas and sugar-corn
afford further instances. Strawberries without runners have come true
from seed ever since their first appearance, over a hundred years ago.
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