hybrid nature, nor due to innate inconstancy of the variety, but must be
considered as pure atavism. I refer to the bud-variations of so many of
our cultivated varieties of shrubs and trees. Many of them are
cultivated because of their foliage. They are propagated by grafting,
and in most cases it is probable that all the numerous specimens [179]
of the same variety have been derived in this way from one primitive,
aberrant individual. We may disregard variegated leaves, spotted or
marked with white or yellow, because they are too inconstant types.
We may next turn our attention to the varieties of trees with cut
leaves, as the oakleaved _Laburnum_, the parsley-leaved vine and the
fern-leaved birch. Here the margin of the leaves is deeply cut and
divided by many incisions, which sometimes change only the outer parts
of the blade, but in other cases may go farther and reach, or nearly
reach, the midvein, and change the simple leaf into a seemingly compound
structure. The anomaly may even lead to the almost complete loss of all
the chorophyll-tissue and the greater part of the lateral veins, as in
the case of the cut-leaved beech or _Fagus sylvatica pectinata_.
Such varieties are often apt to revert by buds to the common forms. The
cut-leaved beech sometimes reverts partially only, and the branches
often display the different forms of cut-leaved, fern-like, oak-leaved
and other variously shaped leaves on the same twigs. But this is merely
due to the wide variability of the degree of fissure and is to be
considered only as a fluctuation between somewhat widely distant
extremes, which may even apparently include [180] the form of the common
beech-leaves. It is not a bud-variation at all, and it is to be met with
quite commonly while the true reversions by buds are very rare and are
of the nature of sports appearing suddenly and remaining constant on the
same twig. Analogous phenomena of wide variability with true reversion
may be seen in the variety of the European hornbeam called _Carpinus
Betulus heterophylla_. The leaves of this tree generally show the
greatest diversity in form. Some other cases have been brought together
by Darwin. In the first place a subvariety of the weeping-willow with
leaves rolled up into a spiral coil. A tree of this kind kept true for
twenty-five years and then threw out a single upright shoot bearing flat
leaves. The barberry (_Berberis_) offers another case; it has a well
known variety with seedless fruit, which can be propagated by cuttings
or layers, but its runners are said always to revert to the common form,
and to produce ordinary berries with seeds. Most of the cases referred
to by Darwin, however, seem to be doubtful and cannot be considered as
true proofs of atavism until more is known about the circumstances under
which they were produced.
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