The name, arm-footed, was given them in reference to a pair of special
structures called the arms, bearing a large number of tentacles; it is
now more frequently spoken of as the lophophore (see p. 122), and
regarded as comparable to the lophophore of the Polyzoa, spread out into
two portions. With the latter group the Brachiopods were formerly united
by Huxley, under the name of Molluscoidea. This name is now obsolete,
because it is understood that all these creatures are widely different
from Molluscs; but the theory of relationship of the Brachiopoda to the
Polyzoa, implied in it, still holds good.
The chief importance of this group lies in its fossil forms, which are
exceedingly numerous, particularly in the Mountain Limestone of the
Carboniferous Period; it is crowded with their shells, especially a form
named, from its elongated shape, Productus. The shells of Brachiopods
are equal-sided; that is to say, the right and left valves match; but
they are inequivalve, the ventral valve being much the biggest. It often
contains a foramen, or hole, at the beak, for the passage of the
pedicle, or stalk, by which the animal is attached to the ground (_e.g._
_Terebratula_, _Rhynchonella_). Sometimes, however, the pedicle passes
out between the valves (_Lingula_, _e.g._), in which case there is no
foramen; or it may be arranged in other ways. Sometimes the shell is
merely attached to the ground by its side, like an oyster. Some forms
are enormously widened in a lateral direction, _e.g._ _Spirifera_, and
the _Productus_ above named. _Lingula_, among others, is remarkable as
being a form that has survived from the earliest geological period to
the present day.
TABLE SHOWING THE CLASSIFICATION OF THE BRACHIOPODA
{ TESTICARDINIS.--Shell Calcareous, with hinges.
{ Skeleton present in the arms.
=BRACHIOPODA.= {
{ ECARDINES.--Shell comparatively soft, composed
{ of Chitin, only strengthened by deposits
{ of lime, without hinges. No skeleton in
{ the arms.
The larva, in its best known forms, passes through the typical larval
stages of the animal kingdom. It is first a one-layered larva, then a
two-layered form, and then becomes a ciliated animal. In this three
regions may be distinguished, representing respectively the head, body,
and pedicle.
The shells of the Brachiopoda, including the kinds above named, may be
seen by the reader in any geological museum.
CHAPTER XI
THE POLYZOA; MOSS-CORALS AND SEA-MATS
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