These queer-shaped things, the Sea-Urchins and their allies, are perhaps
the last creatures amongst which we should think of looking for
relations of the Worms. Yet the earliest stages of the larva are
considered to present a certain amount of resemblance to the Wheel-ball
larva, which has been referred to elsewhere (pp. 42 and 72). Still more
startling fact, these larvæ have been compared to that of
_Balanoglossus_, the lowest member of the Chordata, and a relation of
the Vertebrates themselves (see p. 143).
TABLE SHOWING THE CLASSIFICATION OF THE ECHINODERMATA
{ ECHINOIDEA, OR SEA-URCHINS.
{ ASTEROIDEA, OR STAR-FISHES.
=ECHINODERMATA.= { OPHIUROIDEA, OR BRITTLE-STARS.
{ CRINOIDEA, OR FEATHER-STARS AND
{ STONE-LILIES.
{ HOLOTHUROIDEA, OR SEA-CUCUMBERS.
CHAPTER XIII
THE CHORDATA
The older zoologists used to speak of Vertebrata and Invertebrata as
animals with a back-bone and animals without one, and everyone thought
it a very natural way of dividing up the animal kingdom. It never
occurred to anyone that it was possible to bridge the interval between
them and find a link between the two. But now the Vertebrata have been
compelled to give up their aristocratic pretensions, and own that they
have risen from the ranks of the common people of the animal world; in
other words, that they are descended from the Invertebrates. Their
family secrets have been published to the world, and now everybody knows
that they have poor relations. But how many, and how nearly related?
This we do not accurately know; consequently the whole zoological world
for many years has concentrated all its energies on attempts to find out
the truth about the matter.
A great sensation was caused by the first discovery of a poor relation
of the vertebrates among the Ascidians, or Leather-bottle animals. These
are named from their shape and texture, for they have a leathery skin.
Now some of these Ascidians have larvæ with a tail; and in the tail
there is a long cord-like structure, which in many essential particulars
resembles the cord which precedes the back-bone in the vertebrate
embryo. This structure is called the Notochord (a string down the back).
The credit of this great discovery belongs to Russia; for the presence
of the Notochord in the Ascidian larva was discovered by A. Kowalevsky,
in 1866.
To the present generation of zoological students, the Chordate
affinities of Ascidians are part of the ABC of knowledge; and it is
hardly possible for them to realise that it is only thirty years ago
since the idea was so new that Huxley, in his "Text-book of the
Vertebrata," only alluded to it in a footnote. Would-be zoological
critics, at a somewhat later period, met the theory with ridicule, for
want of better argument. For critics include not only "those who have
failed in literature and art," but also those who have failed in
science.
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