Some of the Ascidians present what has been already described in other
types (p. 57) as "alternation of generations." The discovery of this
fact was made by the poet Adelbert von Chamisso. Some of his verses are
known to English readers, for whom they were translated by Mary Howitt,
a poetess whose writings were popular with our grandmothers, and
deserved to be so. This is not the only case in which a poet has been
also a zoologist: Goethe studied the science, and framed a theory
regarding the vertebrate skull, which he regarded as consisting of a
series of vertebræ. In this he was less fortunate than the Italian
poet; for while Chamisso's observations were correct, and were confirmed
by subsequent writers, Goethe's theory of the skull is anything but
correct. It was made worse, too, by the speculations of subsequent
writers, who attempted to follow it into detail, with the result of
demonstrating its absurdity.
CHAPTER XIV
THE VERTEBRATA
We have spoken of the Notochord as a structure which precedes the
formation of the spinal column in Vertebrates. This needs a little more
definite explanation. We all know that the spinal column of vertebrates
is formed to protect the spinal cord. This protection is, however, an
afterthought, so to speak, of the vertebrate structure; the lowest of
all vertebrates is quite without it; and in the lower groups of fishes
we may trace various steps of its formation. But in these cases where
the spinal column is absent or incomplete, there is a large and
well-developed notochord; and in the embryo of higher vertebrates, when
the spinal column has not yet begun to be formed, the notochord is
equally a conspicuous feature. It runs from the region known as the
mid-brain, to the end of the tail, and lies throughout just beneath the
spinal cord. Whatever its original use in the animal body may have
been, it undoubtedly acts now as a support to the spinal cord, and
indeed to the whole body. Bones, we must explain, do not exist either in
the lower vertebrate, or in the early embryo. In the latter they are
formed by degrees. The spinal cord and the notochord each begin to be
surrounded by rings of cartilage or gristle, which by degrees is changed
into bone. The rings surrounding the notochord, however, gradually
encroach upon it and obliterate it. The place where it has been becomes
the Centrum, or most solid part of each vertebra. The notochord at first
is continuous, and has no division into successive parts; but when the
bony spinal column is developed, it consists of a series of successive
vertebræ. Each of them is made up of several parts, which by degrees
become consolidated into the vertebræ.
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