There is a huge gap between the Lancelet and the true vertebrates. The
lowest form of the latter is _Ammocoetes_, the larva of the Lamprey
(_Petromyzon_). The latter, even in the adult form, has no true limbs,
though there are fringing fins. The notochord sheath is supplemented,
however, by cartilage bars which are equivalent to the beginnings of the
vertebræ of the back-bone. The gills are very different from those of
other true vertebrates, and it has no jaws. Teeth it has, however, on
the tongue and the lining of the mouth. Probably this creature is
greatly altered by adaptation to its peculiar mode of life, so that no
certain conclusions can be drawn from it regarding the structure of
primitive fishes. It has a sucking mouth, by means of which it hangs on
to fishes, while it rasps away their flesh with its rough tongue. When
not thus engaged, it hangs on to a stone by means of its suctional
mouth, thus fixing itself at rest. The Hag-fish, _Myxine_, in many
respects similar, devours dead fishes chiefly. The Hag-fish is found on
English coasts: so is the Marine Lamprey; while two freshwater forms are
found in streams.
Leaving the Cyclostomata, as the above fishes are called, we reach the
true fishes, which have limbs and scales. Something has already been
said regarding their teeth and gills. The Cartilaginous fishes, in
which most part of the skeleton remains gristle and does not become
transformed into bone, include the Sharks, Rays, and Dog-fishes, all
savage animals with strong teeth. The common spotted Dog-fish of our own
shores is familiar to everybody: fishermen regard it with disgust, as it
is not eatable. The Rays are flattened fishes, which live at the bottom
of rather deep water, and attain enormous size even on our own coasts.
The Thornback Skate is covered with prickles (placoid scales). All these
fishes are grouped under the name of Elasmobranchii, the Strap-gilled,
so called from the structure of the gill-arches.
The majority of familiar fishes, such as the herring, mackerel, cod and
sole, belong to the group of _Teleostei_, or Bony Fishes, in which, by
contradistinction from the last group, as much of the skeleton as
possible becomes bone. Nevertheless, traces of the notochord persist in
the back-bone of these fishes. Break the back-bone across, of a cod or a
sole, and you will find between adjacent sides of the centra, or middle
parts of the vertebræ, a pad of gristly substance. This is the remaining
substance of the notochord, which finds room between the cup-shaped
sides of the centra. When the centrum, instead of being biconcave, is
solid, as in the higher Vertebrata, the notochord is obliterated by its
encroachment.
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