The following considerations, which submit themselves with reference to
seasonal dimorphism, are readily conceivable, at least, in so far as
they can be arrived at by purely theoretical methods. Were the summer
to become shorter the generation which formerly hibernated in the pupal
stage would be advanced further into the spring. It would not thereby
necessarily immediately lose the winter characters which it formerly
possessed. Whether this would happen, and to what extent, would rather
depend upon the intensity of the action of the summer climate on the
generation in question, and on the number of generations which have
been submitted to this action. Hitherto no attempts have been made to
expose a monomorphic species to an elevated temperature throughout
several generations, so as to obtain an approximate measure of the
rapidity with which such climatic influences can bring about changes.
For this reason we must for the present refrain from all hypothesis
relating to this subject.
The disturbance of generations by the shortening of summer might also
occur to a species in such a manner that the generation which formerly
hibernated advances into the spring, the last of the summer generations
at the same time reaching the beginning of winter. The latter would
then hibernate in the pupal state, and would sooner or later also
assume the winter form through the action of the cold of winter. We
should, in this case, have two generations possessing more or less
completely the winter form, the ancient winter generation now gradually
losing the winter characters, and the new winter generation gradually
acquiring these characters.
In the reverse case, _i.e._, by a lengthening of the summer, we
should have the same possibilities only with the difference that the
disturbance of generations would occur in a reverse direction. In this
case it might happen that the former winter generation would become the
autumnal brood, and more or less preserve its characters for a long
period. Here also a new winter generation would be produced as soon as
the former spring brood had so far retrograded that its pupæ hibernated.
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