It is erroneous to assert that Classification has only to take
form-relationship into consideration; that it should and can be
nothing else than the expression of form-relationship. The latter
is certainly our only measure of blood-relationship, but those who
maintain the assertion that form- and blood-relationship are by no
means always synonymous, are undoubtedly correct. I shall in a future
essay adduce facts which leave no doubt on this point, and which prove
at the same time that modern systematists--especially in the order
Lepidoptera--have always endeavoured--although quite unconsciously--to
make the blood-relationship the basis of their classification. For
this reason alone, larvæ and pupæ would have an important bearing upon
the establishment of systematic groups, although certainly in a manner
frequently irregular.
It must be admitted that so long as we are able to compare the species
of one group with those of another _in one form only_, we are often
unable to ascertain the blood-relationship.[68] In such cases we can
only determine the latter from the form-relationship, and as these
are not always parallel, any conclusion based on a single form must
be very unsound. If, for instance, butterflies emerged from the egg
directly, without passing through any larval stage, a comparison of
their resemblances of form would alone be of systematic value; we
should unite them into groups on the ground of these resemblances
only, and the formation of these groups would then much depend upon
the weight assigned to this or that character. We might thus fall into
error, not only through a different valuation of characters but still
more because two species of near blood-relationship frequently differ
from one another in form to a greater extent than from other species.
We should have no warrant that our conception of the form-relationship
expressed the genealogical connection of the species. But it would
be quite different if every species presented itself in two or three
different forms. If in two species or genera the butterflies as well
as the larvæ and pupæ exhibited the same degree of form-relationship,
the probability that this expressed also the blood-relationship would
then be exceedingly great. Now this agreement certainly does not always
occur, and when these different stages are related in form in unequal
degrees, the problem then is to decide which of these relationships
expresses the genealogy. This decision may be difficult to arrive at
in single cases, since the caterpillar may diverge in form from the
next blood-related species to a greater extent than the butterfly, or,
conversely, the butterfly may diverge more widely from its nearest
blood-related species than the caterpillar.
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