I have been able to compare altogether about forty caterpillars,
thirty-five of which were living. All these specimens possessed nearly
the same greyish-green ground-colour, and most of them had exactly
the simple marking as represented, for instance, in Hübner’s figure,
_i.e._, a rather broad greenish-white subdorsal line, somewhat faded
at the edges, and without a trace of spots on any of the segments
with the exception of the eleventh, on which there was a yellowish,
black-bordered mirror-spot, with a broad, diffused, vivid orange-red
nucleus. Specimens also occur, and by no means uncommonly, in which no
other markings are to be seen than those mentioned; there were nine
among twenty-eight examples compared from this point of view.
In many other individuals of this species small red spots appear on
the subdorsal line, exactly in the positions where the ring-spots are
situated in the other species of the genus (Fig. 60), so that these
spots are thus repetitions of the single ring-spot--a fact which must
appear of the greatest interest in connection with the development
of the markings throughout the whole genus. But this is not all, for
again in other specimens, these red spots stand on a large yellow
“mirror,” and in one individual (Fig. 59), they had become developed
into well-formed ring-spots through the addition of a black border. We
have thus presented to us in one and the same stage of a species, the
complete development of ring-spots from a subdorsal line.
These facts acquire a still greater interest, as showing how new
elements of marking are produced. The spots on the subdorsal line
decrease from the posterior to the anterior segments, so that they
must undoubtedly be regarded as a repetition or transference of the
ring-spot previously developed on the eleventh segment. I will now
proceed to furnish proofs in support of this statement.
I have never met with any specimens having ring-spots on all the
segments--in the most prominent instances these spots were present
on segments 10-5. This was the case in three out of the twenty-eight
caterpillars minutely examined. On all these segments, however, the
ring-spots were not equally developed, but increased in perfection
from the posterior towards the anterior segments. In the larva
represented in Fig. 59 for example, there is a completely developed
ring-spot on segment 10, which, although possessing but a feeble black
“ground-area,” is still distinctly bordered; on segment 9 this border
is less sharp, and not so dark, and it is still less sharp and much
lighter on segments 8 and 7, whilst it has completely disappeared from
segment 6, the yellow “mirror” having at the same time lost in size. On
segment 5, only two small contiguous reddish spots, the first rudiments
of the nucleus,[94] can be recognized on close inspection.
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