The explanation of such a case--_i.e._ its reference to known causes of
species transformation--is never easy, and is indeed impossible without
a precise knowledge of the ontogeny of many species, as well as of the
original significance of the characters in question. In the case of the
_Deilephila_ larvæ, however, such knowledge is still wanting. It is
true that they present us with parallel developmental series, but these
do not depend on an unknown phyletic force--the parallelism can be
referred to the action of the imperfectly known laws of growth innate
in segmented organisms. Because the characters of one segment have
a tendency to repeat themselves on the others, from one parent-form
possessing ring-spots on one segment only, there may have proceeded
several developmental series, all of which developed rows of such spots
independently of each other.
From these considerations we may venture to construct the following
genealogical tree:--
[Illustration: POSSIBLE GENEALOGICAL TREE OF THE GENUS DEILEPHILA
The circles indicate the phyletic stages IV.-VIII.; the eighth is only
reached by _Nicæa_, and is distinguished from the seventh chiefly by
the ontogeny, in the third stage of which the seventh phyletic stage is
reached, whilst in _Euphorbiæ_ and _Dahlii_ this stage is reached in
the fourth ontogenetic stage. The phyletic stages indicated by queries
are extinct, and only known through the ontogeny of existing species.
It must be understood that this pedigree expresses only the ideal and
not the actual relations of the species to one another. Thus, it is
possible that _Hippophaës_ is not the parent-form, but an unknown or
extinct species, which must, however, have possessed the same marking,
and so on.]
Four parallel series here proceed from the parent-form _Hippophaës_;
there may have been five, or possibly only three, but the incomplete
state of our knowledge of the ontogeny does not permit of any certain
conclusion. For the point under consideration this is, however, quite
immaterial. The distance from the central point (the parent-form)
indicates the grade of phyletic development which the respective
species have at present reached.
There is another case which is no less instructive, because it reveals,
although in a somewhat different manner, the action of a law of growth
innate in the organism itself, but which can nevertheless by no means
be regarded as equivalent to a phyletic vital force. I refer to the
coloured edges of the oblique stripes which occur in most of the
species of the genus _Sphinx_. It has already been insisted upon in
a previous section, that the mode in which this character originates
negatives the assumption of a phyletic force, because these coloured
edges are gradually built up out of irregularly scattered spots. There
is no occasion for a “developmental force” to grope in the dark; if
such a power exists, we should expect that it would add new characters
to old ones with the precision of a master workman.
Public-domain text, read in full here on John Shaqi.
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