insignificant black spots on the under side of the wing (_P. Alexis_
female, and _P. Agestis_); and this diagnosis proves sufficient, since
_P. Alexis_, which has the spots in a straight row, has a different
caterpillar to _P. Agestis_, in which the central spot is nearer the
base of the hind wing!
For the reasons just given, I maintain that it is neither justifiable
nor useful to designate the di- and polymorphism of butterflies as
di- and polychroism, and thereby to attribute but little importance to
these phenomena.[44] This designation would be only justifiable if the
differences of colour were due to other causes than the differences of
form, using this last word in a narrow sense. But it has been shown
that the same direct action of climate which originates new colours,
produces also in some species differences of form (contour of wing,
size, &c.); whilst, on the other hand, it has long been known that many
protective colours can only be explained by the indirect action of
external conditions.
When I raise a distinction in the nature of the changes between
seasonal dimorphism and the remaining known cases of heterogenesis,
this must be taken as referring only to the biological or physiological
result of the change in the transformed organism itself. In seasonal
dimorphism only insignificant characters become prominently changed,
characters which are without importance for the welfare of the species;
while in true heterogenesis we are compelled to admit that useful
changes, or adaptations, have occurred.
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