On the other hand, it is equally conceivable that heterogenesis may
have been developed from true metagenesis in the case of larvæ which,
having acquired the faculty of asexual propagation, are similar in
function to sexually mature insects. This possibility is not at first
sight apparent. If the nursing-larvæ of the _Cecidomyiæ_ were as much
like the sexual insects as are the young Orthoptera to the sexually
mature forms, we should not know whether to regard them as degraded
sexual insects, or as true larvæ which had attained the power of
asexual propagation. Their propagation would be considered to be
parthenogenesis; and as it could not be denied that heterogenesis was
here manifest, the mode of development of their particular kind of
propagation might be proved, i.e., it might be demonstrated, that the
generations now parthenogenetic were formerly mere reproductive larval
stages.
I have only offered these last observations in order to show on what
uncertain ground we are still standing with regard to this subject
whenever we deal with the meaning of any particular case, and how
much still remains to be done. It appears certain that the two forms
of cyclical propagation, heterogenesis and metagenesis, originate in
entirely distinct ways, so that it must be admitted that, under these
circumstances, the idea of the existing conditions respecting the true
genesis may possibly be erroneous. To indicate the manner in which the
cyclical mode of propagation has arisen in any single case, would only
be possible by a searching proof and complete knowledge of existing
facts in addition to experiments.
VI.
GENERAL CONCLUSIONS.
I shall not here give a repetition and summary of the results arrived
at with respect to seasonal dimorphism, but rather the general
conclusions derived from these results; and, at the same time, I may
take the opportunity of raising certain questions which have not
hitherto found expression, or have been but briefly and casually stated.
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