Herein might be sought the explanation of the different strength with
which the reproductive power is affected; not indeed in the direction
of the leap itself, but in the differences in the ontogeny which
are determined by the differences in the direction of the leap. The
reversion of the Triton to an older phyletic stage coincides with
the arrest at a younger ontogenetic stage; or, in other words, the
older stage of the phylogeny to which reversion takes place is still
entirely comprised in the ontogeny of each individual. Each Triton is
perennibranchiate throughout a long period of its life; the reverting
individual simply reverts to the older phyletic stage by remaining at
the larval stage of its individual development.
But it is quite different with the reversion of the Axolotl to the
formerly acquired, but long since abandoned Amblystoma form. This is
not retained in the ontogeny of Axolotl, but has been completely lost;
for a long series of generations--so must we suppose--the ontogeny
has always only attained to the perennibranchiate form. Now if at the
present time certain individuals were compelled to revert to the
Amblystoma form, certainly no greater leap would have been made from
a morphological point of view, than in the reversion of Triton to the
perennibranchiate form, but at the same time the leap would be in
another direction, viz. over a long series of generations back to a
form which the species had not produced for a long period, and which
had to a certain extent become foreign to it. We should thus have here
also the grafting of a widely different constitution upon that of the
Axolotl, or, if one prefers it, the commingling of two widely different
constitutions.
Of course I am far from wishing to pretend that this “explanation” is
exact; it is nothing more than an attempt to point out the direction
in which the causes affecting the reproductive powers in different
degrees are to be looked for. A deeper penetration into and special
demonstration of the manner in which these causes bring about such
results, must be reserved for a future period. For the present it
must suffice to have indicated that there is an essential distinction
between the two kinds of reversion, and to have made it to some
extent comprehensible that this distinction may be the determining
impulse with respect to the question of sterility. Perhaps the law
here concealed from us may one day be thus formulated:--Atavistic
individuals lose the power of reproduction the more completely, the
greater the number of generations of their ancestors whose ontogeny no
longer comprises the phyletically older stage to which the reversion
takes place.
Public-domain text, read in full here on John Shaqi.
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