Now according to this law, each step in phyletic development when
replaced by a later one, must remain preserved in the ontogeny, and
must therefore appear at the present time as an ontogenetic stage
in the development of each individual. But my interpretation of the
transformation of the Axolotl appears to stand in contradiction to
this, since the Axolotl, which at a former period was an Amblystoma,
retains nothing of the latter in its ontogeny. The contradiction is,
however, only apparent. As long as we are concerned with an actual
advance in development, and therefore with the attainment of a new step
never formerly reached, the older stages will be found in the ontogeny.
But this is not the case when the new stage is not an actual novelty,
but formerly represented the final stage of the individual development;
or, in other words, when we are concerned with the reversion, not
of single individuals, but of the species as such, to the preceding
phyletic stage, _i.e._ with a phyletic degeneration of the species.
In this case the former end-stage of the ontogeny would be simply
eliminated, and we should then only be able to recognize its former
existence by its occasional appearance in a reversion form. Thus, under
certain conditions the Triton sinks back to the perennibranchiate
stage; not in such a manner that the individual first becomes a Triton
and then undergoes perennibranchiate re-modification, but simply, as I
have already shown above, by its remaining at the Ichthyodeous stage
and no longer attaining to the Salamander form. So also, according
to my hypothesis, the salamandrine _Amblystoma Mexicanum_, formerly
inhabiting the shores of the Lake of Mexico, has degenerated to the
perennibranchiate stage, and the only trace that remains to us of its
former developmental status is the tendency, more or less retained
in each individual, to again ascend to the salamander stage under
favourable conditions.
The third and last consequence which my interpretation of the facts
entails, is the change in the part played by reversion in organic
nature. Whilst atavistic forms have hitherto been known only as
isolated and exceptional cases, interesting indeed in the highest
degree, but devoid of significance in the course of the development
of organic nature, a real importance in this last respect must now be
attached to them.
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