One might indeed insist that with us also the land-salamanders conceal
themselves in moist places during dry weather, and often lie hidden,
as in Mexico, in a hole, in a cluster of as many as ten together;
but with us they leave their lurking-place from time to time and go
in search of food. Señor Velasco mentions nothing with respect to
this. What especially struck me was the statement that the Mexican
Amblystomas _were also to be found in the water_.[281] When Lake Santa
Isabel is drained, the fishermen stretch large nets across the exit
channels, and in these they not only find ordinary Axolotls, but also
some “_sin aretes_,” which they also designate “_mochos_,” _i.e._
hornless Axolotls, because they have no gills, but have already reached
the Amblystoma stage. Our land-salamanders live in the water only
as larvæ, but they also love and require moisture. Only the female
enters the water when she wants to deposit her young (eggs with mature
larvæ), and then only at the margin of shallow pools or small brooks.
The Mexican Amblystoma thus much more resembles in its habits our
water-salamanders (Tritons), which remain in the water at least during
the whole period of reproduction. These also leave the water later,
and, like the land-salamander, seek concealment in the earth. They have
this habit also in those districts which possess a very dry atmosphere;
and especially in the Engadine, where I first conceived the idea of
taking into account the dryness of the air, I found in the pools at the
end of August and the beginning of September only larvæ of Tritons. The
older Amphibians must therefore have been on the land, presumably in
their places of winter concealment.
From what we have hitherto learnt from Señor Velasco, the mode of
life of _Amblystoma Tigrinum_ must resemble that of our Tritons,
although its structure is that of a land-salamander. I would thus
offer the following explanation of the facts at present known:--Owing
to the periodic drying up of the lake of Santa Isabel, the _Siredon
Tigrinus_ would be again compelled to undergo metamorphosis. Whether
this was formerly entirely abandoned, or whether it always occurred
in solitary individuals, is almost immaterial; in any case the habit
of metamorphosis must have been very rapidly acquired through natural
selection, and must have again become general, if the faculty was only
present in the species, although latent. Through the dryness of the
air, the Amblystomas that had taken to the land would be compelled to
bury themselves at once, and to remain asleep till the recurrence of
the rainy season, when they would hasten back into the water and would
there live as a species of Triton.
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