In the second essay the attempt was next made with reference to double
form-relationship, as presented for observation in metamorphic insects,
to draw conclusions as to the causes of the transformations. It
appeared here that form- and blood-relationship do not always coincide,
since the larvæ of a species, genus, or family, &c., may show quite
different form-relationships to their imagines. These facts alone told
very decisively against the existence of an internal developmental
power, so that the latter had likewise to be set aside by the method
of elimination, since the observed incongruences as well as the
congruences of form-relationship, found sufficient explanation in the
action of the environment on the organism.
This investigation thus also led to the denial of a phyletic vital
force.
In the third essay I finally sought to prove that the only case
of transformation of one species into another at present actually
observed[282], could not without further evidence be interpreted as
the result of the action of a phyletic vital force, but that more
probably we had here only an apparent case of new formation, which was
in reality but a reversion to a stage formerly in existence.
If this last investigation removes the only certain observation which
could have been adduced in favour of the hypothesis of a phyletic vital
force, so also do the two former essays show that this hypothesis, at
least in the case of insects, must be abandoned as inadequate.
The question now arises whether this conclusion, based on such a
limited range of inquiry, can also be applied to the other groups of
the organic world without further evidence.
The supporters of a principle of organic development will deny this in
each individual case, and will demand special proof for each group of
organisms; I believe this position, however, to be incorrect. Here,
if anywhere, it appears to me justifiable to apply the conclusions
inductively from special cases to general ones, since I cannot at all
see why a power of such pre-eminent and fundamental importance as a
phyletic vital force should have its activity limited to solitary
groups in the organic world. If such a power exists it must be the
inciting cause of organic development in general, and must be equally
necessary in every part of creation, as no advancement could take place
without it. In this case, however, the force would be recognizable and
demonstrable at every point; the phenomena should nowhere stand in
opposition to its admission, and should in no case be explicable or
comprehensible without it. The same laws and forces which caused the
development of one group of forms must underlie the development of the
whole organic world.
I therefore believe that we are correct in applying to the whole living
world the results furnished by the investigation of insects, and in
thus denying the existence of an innate metaphysical developmental
force.
Public-domain text, read in full here on John Shaqi.
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