I will first answer the second question. On all possible sides the
earlier view of the absolute nature of species is contradicted; there
is no boundary between species and varieties. But when Von Hartmann
assumes that by the transformation of one species “into another” the
“whole uniformly connected complex must become changed,” he falls
back into the old doctrine of the absolute nature of species, which
is sharply contradicted by multitudes of facts. We not unfrequently
observe varieties which differ from the parent-form by only a single
character, whilst others show numerous differences, and again others
may be seen in which the differences predominate. This last deviation
would then be designated by many systematists as a new species, but not
so by others.
The “specific type” is thus indeed a kind of mosaic-work, but it is
a structure to which all the single characters--the stones of the
mosaic--belong and build up one harmonious whole, and not a meaningless
confusion. Some of the stones or groups of stones can be taken away
and replaced by others differently coloured without the structure
being thereby necessarily distorted, _i.e._ destroyed as a structure;
but the larger the stones which are exchanged the more necessary will
corrections in the other parts of the structure become, in order that
the harmony of the whole may be preserved.
Still more weighty than those insensible transitions which in various
groups of animals so frequently connect species with species, appear
to me, however, the facts made known in the second essay of the
second part of this volume, which prove that the two forms in which
one species appears can change entirely independently of one another.
The caterpillar changes and becomes a new variety or even species
(according to the form-value of the change), whilst the butterfly
remains unaltered. How could this occur if some other law than that
of physiological equilibrium linked together the parts or characters
and permitted them to become severed? Must not the two stages become
changed with and through one another, like the parts of one body, since
they first together constitute the specific type? Is not the fact of
this not happening a proof that the whole “uniformly connected complex”
of the specific type is not bound and held together by a metaphysical
principle, but simply by natural laws?
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