This form of incongruence is always connected with unequal divergence
between the two stages of the one systematic group--in this
case the _Terebrantia_. The larvæ of this imaginal group partly
possess caterpillar-like (_Phytospheces_) and partly maggot-formed
(_Entomospheces_) larvæ, and differ from one another to a considerably
greater extent than the saw-flies from the ichneumons.[211] The final
cause of the incongruence lies therefore in this case also in the fact
that one stage has suffered stronger changes than the other, so that
a deeper division of the group has occurred in the former than in the
latter.
The analogous incongruences in single families of the Lepidoptera may
have arisen in a similar manner, as has already been more clearly shown
above; only in these cases we are as yet unable to prove in detail that
the larval structure has become more strongly changed through special
external conditions of life than that of the imagines.
In the smallest systematic group--varieties, it has been possible to
furnish some proof of this. The one-sided change here depends in part
upon the _direct action_ of external influences (seasonal dimorphism,
climatic variation), and it can be shown that these influences
(temperature) acted only on the one stage, and accordingly induced
change in this alone whilst the other stage remained unaltered.
It has now been shown--not indeed in every individual case, but for
each of the different kinds of incongruence of form-relationship--that
there is an exact parallelism corresponding throughout with the
incongruence in the conditions of life. Wherever the forms diverge
more widely in one stage than in the other we also find more widely
divergent conditions of life; wherever the morphological systemy of
one stage fails to coincide with that of the other--whether in the
extent or in the value of the groups--the conditions of life in that
stage also diverge, either more widely or at the same time within
other limits; whenever a morphological group can be constructed from
one stage but not from the other, we find that this stage alone is
submitted to certain common conditions of life which fail in the other
stage.
The law that the divergence in form always corresponds exactly with
the divergence in the conditions of life[212] has accordingly received
confirmation in all cases where we have been able to pronounce
judgment. Unequal form-divergences correspond precisely with unequal
divergence in the conditions of life, and community of form appears
within exactly the same limits as community in the conditions of life.
These investigations may thus be concluded with the following law:--In
types of similar origin, _i.e._ having the same blood-relationship,
the degree of morphological relationship corresponds exactly with the
degree of difference in the conditions of life in the two stages.
With respect to the question as to the final cause of transformation
this result is certainly of the greatest importance.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account