A complete parallel can be drawn between the two classes of
developmental phenomena. From the very different systematic values
attached by taxonomists to this or that organ in a group of animals,
it may be concluded that the individual parts of an organism are to
a certain extent independent, and that each can vary independently,
when affected either entirely alone or in a preponderating degree by
transforming impulses, without all the other parts of the organism
likewise suffering transformation, or at least without their becoming
modified in an equal degree. Did all the parts and organs in two groups
of animals diverge from each other to the same extent, the systematic
value of such parts would be perfectly equal; we should, for example,
be able to distinguish and characterize two genera of the family of
mice by their kidneys, their liver, their salivary glands, or by the
histological structure of their hair or muscles, or even by differences
in their myology, &c. equally as well as by their teeth, length of
toes, &c. It is true that such a diagnosis has yet to be attempted; but
it may safely be predicted that it would not succeed. Judging from all
the facts at present before us, the individual parts--and especially
those connected in their physiological action, _i.e._ the system of
organs--do not keep pace with reference to the modifications which the
species undergoes in the course of time; at one period one system and
at another period some other system of organs advances while the others
remain behind.
This corresponds exactly with the result already deduced from the
unparallel development of the independent ontogenetic stages. If the
inequality in the phyletic development is more sharply pronounced in
this than in the last class of cases, this can be explained by the
greater degree of correlation which exists between the individual
systems of organs in any single organism as compared with that existing
between the ontogenetic stages, which, although developed from one
another, are nevertheless almost completely independent. We should have
expected _à priori_ that a strong correlation would have here existed,
but as a matter of fact this is not the case, or is so only in a very
small degree.
Just as in the stages of metamorphosis the inequality of phyletic
development becomes the more obliterated the more distant and
comprehensive, or, in other words, the greater the period of existence
of the groups which we compare, so does the unequal divergence of the
systems of organs become obliterated as we bring into comparison larger
and larger systematic groups.
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