In this fourth stage the larva of _D. Hippophaës_ is of a lighter green
(Fig. 58), the subdorsal yellowish with sharp boundaries, and the
infra-spiracular line pure white, as in the next stage. The shagreening
is present, but none of the shagreen-dots are red or reddish, and no
trace of a ring-spot can be detected on the subdorsal line with the
exception of that on the eleventh segment. In this last position this
line is somewhat widened, and a long, diffused, rose-red spot can there
be recognized upon it (Fig. 58 A). The black “ground-area” present
in the fifth stage is as yet absent, and the spot is not so sharply
separated anteriorly from the subdorsal line as it becomes later.
From these observations we might venture to expect that in the third
stage of _Hippophaës_, the subdorsal line would also be free from this
spot on the eleventh segment, and it is possible that in the second
stage this line is itself absent.
THE GENUS DEILEPHILA: SUMMARY OF FACTS AND CONCLUSIONS.
Regarding only the _adult_ larvæ of the species of _Deilephila_,
these represent in their five groups, five stages in the phyletic
development of the genus; but if we also take into consideration the
developmental history, two more stages must be added, viz., that in
which the caterpillar possesses no particular marking, as was found to
be the case in the first stage of the development of _D. Euphorbiæ_
and _D. Dahlii_; and a second stage with a subdorsal line, but without
any ring-spot formations. Seven stages of phyletic development must
therefore be distinguished.
_Stage 1._--No species with entire absence of marking in the adult form
now occurs.
_Stage 2._--A subdorsal, accompanied by a spiracular line, extends from
the caudal horn to the first segment. This also no longer forms the
final stage of the ontogeny, but is, however, undoubtedly retained in
the second stage of several species (_D. Vespertilio_, _Livornica_,
_Lineata_, and perhaps also _Galii_).
_Stage 3._--The subdorsal line bears a ring-spot on the penultimate
segment; the other markings as in the last stage. _D. Hippophaës_ only
belongs to this stage, a small number of specimens, however, showing
a transition to the following stage by the transference of ring-spots
from the posterior to the anterior segments.
_Stage 4._--Open ring-spots appear on the subdorsal line on all the
segments from the eleventh to the first. _D. Zygophylli_ and the North
American _D. Lineata_ belong here.
_Stage 5._--Closed ring-spots are situated on the subdorsal line. Of
the known species, only _D. Livornica_ concludes its development at
this phyletic stage.
_Stage 6._--A single row of ring-spots replaces the subdorsal line. _D.
Galii_, _Vespertilio_, and _Mauritanica_ represent this stage at the
conclusion of their ontogeny.[95]
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