(2.) There are species showing a high degree of variability in all
three stages. This case must be of rare occurrence, as I am only able
to adduce _Araschnia Prorsa-Levana_, a fact which arises from the
circumstance that the pupal stage is, as a rule, but seldom variable.
(3.) There are species which are variable in two stages and constant
in the third. To this class, for example, belongs _Smerinthus Tiliæ_,
of which the larva and imago are very variable, whilst the pupa is
quite constant. The same is the case with _Lasiocampa Pini_, the
well-known fir moth. Many butterflies show this same phenomenon in
other combinations, such, for instance, as _Vanessa Urticæ_ and
_Polychloros_, in which the larva and pupa are very variable, and the
imago very constant. In a less degree the same is also the case with
_Vanessa Atalanta_, whilst in _Pieris Napi_ the pupa and imago are
variable, and the caterpillar remarkably constant, this likewise being
the case with the local form _Bryoniæ_, which, according to my theory,
is to be regarded as the parent form of _Napi_ (See Part I. of the
present volume).
(4.) There are species which are constant in two stages, and variable
only in the third. Thus, a few species can be found in which the
larva and pupa are constant and the imago variable. This is the case
with _Saturnia Yamamai_, the imago of which is well known to present
numberless shades of colour, varying from light yellow to greyish
black, whilst the green caterpillar shows only slight individual
differences of marking, and scarcely any differences of colour. The
pupa of this species is quite constant. _Arctia Caja_ and _Hebe_, and
_Chelonia Plantaginis_ belong to this same category.
There are a very large number of species which possess very constant
imagines and pupæ, but extremely variable larvæ. The following are
the cases known to me:--_Macroglossa Stellatarum_, _Fuciformis_
and _Bombyliformis_; _Chærocampa Elpenor_, _Celerio_, and _Nerii_;
_Deilephila Galii_, _Livornica_, Hübn., _Hippophaës_, _Vespertilio_,
and _Zygophylli_; _Sphinx Convolvuli_; _Acherontia Atropos_;
_Smerinthus Ocellatus_ and _Tiliæ_; _Callimorpha Hera_; _Cucullia
Verbasci_ and _Scrophulariæ_.
Cases in which the variability depends entirely upon the pupa, while
the larva and imago are extremely constant, are of great rarity.
_Vanessa Io_ is a case in point, the pupa being light or dark brown, or
bright golden green, whilst in the two other stages scarcely any light
shades of colour or variations in the very complicated marking are to
be met with.
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