In seasonal dimorphism, as I have attempted to show, it is the
direct action of climate, and indeed chiefly that of temperature,
which brings about the change in some of the generations. Since
these generations have been exposed to the alternating influence
of the summer and winter temperature a periodical dimorphism has
been developed--a regular cycle of dissimilar generations. It has
already been asserted that the consecutive generations of a species
comport themselves with respect to heredity in a manner precisely
similar to that of the ontogenetic stages, and at the same time such
succeeding generations point out the parallelism between metamorphosis
and heterogenesis. If influences capable of directly or indirectly
producing changes operate on any particular stage of development,
these changes are always transmitted to the same stage. Upon this
metamorphosis depends. In a precisely similar manner changes which
operated periodically on certain generations (1, 3, 5, for instance)
are transmitted to these generations only, and not to the intermediate
ones. Upon this depends heterogenesis. We have just been led to the
comprehension of heterogenesis by cyclical heredity, by the fact
that a cycle is produced whenever a series of generations exists
under regularly alternating influences. In this cycle newly acquired
changes, however minute in character at first, are only transmitted to
a later, and not to the succeeding generation, appearing only in the
one corresponding, i.e. in that generation which exists under similar
transforming influences. Nothing can more clearly show the extreme
importance which the conditions of life must have upon the formation
and further development of species than this fact. At the same time
nothing shows better that the action of these conditions is not
suddenly and violently exerted, but that it rather takes place by small
and slow operations. In these cases the long-continued accumulation
of imperceptibly small variations proves to be the magic means by
which the forms of the organic world are so powerfully moulded. By the
application of even the greatest warmth nobody would be able to change
the winter form of _A. Levana_ into the summer form; nevertheless, the
summer warmth, acting regularly on the second and third generations
of the year, has, in the course of a lengthened period, stamped these
two generations with a new form without the first generation being
thereby changed. In the same region two different climatic varieties
have been produced (just as in the majority of cases climatic varieties
occur only in separate regions) which alternate with each other, and
thus give rise to a cycle of which each generation propagates itself
sexually.
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