Nevertheless, there are many birds that seem to rely entirely upon the
territory to supply them with all that is necessary. Each Warbler seeks
its food within the precincts of its own particular domain, and, except
in occasional instances, neither resorts to neutral ground nor makes
excursions into the locality immediately surrounding the territory, as
does the Bunting. Probably it would be disastrous if it attempted to do
so, for since its young at birth are so delicate and so susceptible to
changes of temperature, it cannot afford to be absent from them for
long. Of the two experiments made with young Whitethroats, one was made
under favourable and the other under unfavourable conditions. In this
latter case the temperature was 50° F., and the young, it may be
remembered, only survived for a little over one hour. Now exposure at
that temperature is evidently dangerous, but it would be still more
dangerous if the weather were wet instead of dry, and the temperature
46° F. instead of 50° F.; and it is, I imagine, on this account that the
impulse to brood is so strongly implanted in the female. No sooner, it
seems, does she depart than she returns with a small quantity of food
which she hurriedly distributes and immediately settles down to brood;
and if forcibly prevented from returning, her attitude betrays symptoms
of what, humanly speaking, we should term great distress. If, then, the
conditions in the external environment were such as would make it
difficult for the female to obtain food rapidly, what advantage would
she derive from so strongly developed an impulse? Might it not be a
disadvantage? Might it not mean that she would abandon the search too
readily and be content to return with an insufficient supply, and might
not that be as injurious to the young as prolonged exposure? Manifestly
the impulse to brood could only have developed strength in so far as it
fitted in with all the other factors that make for survival; and the
principal factor in the external environment seems to be the territory.
How could the young have been freed from the risk of exposure if the
impulse to brood had not been so strongly implanted in the parent? How
could the impulse to brood have been free to develop if a supply of food
had not been first insured? How could the supply of food have been
insured if numbers of the same species had been allowed to breed in
close proximity?
From the foregoing facts it is clear that the young of many species are
at birth susceptible to cold and unable to withstand prolonged exposure.
The parents must therefore be in a position to obtain food rapidly, and
consequently it is important that there should be an ample supply in the
vicinity of the nest. This end the territory certainly serves to
promote; it roughly insures that the bird population of a given area is
in proportion to the available means of subsistence, and it thus reduces
the risk of prolonged exposure to which the young are always liable.
Public-domain text, read in full here on John Shaqi.
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