How, then, can the theory apply to a species that breeds under such
conditions? What part can the territory play in furthering the life of
the individual when large numbers of nests are built closely together in
the same tree? There is much evidence to show that mutual protection is
a necessary condition of the Rook's existence; many cases are on record
of rookeries being destroyed by Carrion-Crows, Hooded Crows, and Ravens.
For instance, Mr. Ward Fowler records a case in which a pair of Crows
attacked a small rookery, ransacked the nests, and destroyed the eggs,
with the result that not a single pair of Rooks was left in the
settlement. Each Rook must therefore secure a position within the
precincts of the community if it is to have a chance of success in the
attainment of reproduction. But every locality cannot supply sufficient
trees of the right kind, appropriately situated and in suitable relation
to the food supply, in which numbers of nests can be built in close
proximity; so that if more than one community were to attempt to
establish itself in a limited area, the supply of food or the supply of
trees might become a pressing problem. Each community must therefore be
prepared to defend its own interests, and each must be regarded as one
unit and the area occupied as one territory within which are included a
number of lesser territories. The individual may fail to establish
itself within a community, but, even if it succeeds, the community may
fail to establish the rights of communal ownership; hence it has to face
a twofold possibility of failure, and if it lacked the inherited nature
which leads the Guillemot to secure a position upon the ledge, or the
Bunting to obtain a position in the marsh, the chances are that it would
fail in the attainment of reproduction.
The question now arises as to how it comes about that the area occupied
by each individual conforms in broad outline to that which has proved
beneficial for the welfare of the species as a whole. We shall find that
up to a point the answer is a simple one. No one could study the
behaviour of animals without observing the important part that habit
plays in the life of the individual; an action performed to-day is
liable to be repeated to-morrow and the following day until it becomes
ingrained in the life of the individual. This must not be taken to mean,
however, that a particular action has to be performed for many days in
succession before it becomes definitely fixed; if only it is repeated a
number of times, even within the space of a few hours, it will acquire
sufficient strength for its continuance; but continued repetition gives
increased fixity, and, as time goes by, it becomes increasingly
difficult for the creature to make a change unless the character of the
situation necessitates readjustment.
Public-domain text, read in full here on John Shaqi.
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