First, then, there is the general consideration, namely, that the enmity
occurs for the most part just at the time when the territories are in
process of being established. During autumn and winter, many birds of
more or less close affinity assemble together in flocks, wherever the
supply of food is abundant, and are then not only sociable, but, so
there is reason to believe, are mutually helpful both in discovering the
necessary means of subsistence which are often none too plentiful, and
in affording protection from enemies, which, on the contrary, are often
numerous. That the different units of which these flocks are composed
should live on amicable terms is therefore as necessary for the welfare
of the whole community at this particular season as that the different
individuals of the same species should do so. But just as the sociable
relations, which obtain between these individuals throughout the winter,
undergo a marked change at the commencement of the breeding season, so,
too, do different species, which habitually associate together, suddenly
become hostile to one another. This change is coincident in time with
the rise of the organic condition which leads to the establishment of
territories; and the hostility continues, though in diminishing degree,
throughout the breeding season, and dies away the following autumn.
For example, different Warblers resort to the elders (_Sambucus nigra_)
in September, and there pass much time feeding on the fruit which is
then ripe and often abundant. In the same bush there may be Blackcaps,
Garden-Warblers, Whitethroats, and Lesser Whitethroats, some preening
their feathers, others searching for the berries, others again, with
feathers relaxed, making feeble attempts to sing. Occasionally there may
be a scuffle, perhaps between a Blackcap and a Lesser Whitethroat, or
between a Garden-Warbler and a Blackcap, but it is of short duration and
lacks vigour. Apart, however, from such temporary disturbances, there is
no real rupture in their relations, and certainly nothing to lead one to
suppose that the bickerings are determined by the functioning of any
specific instinct. Yet only a few months previously some of them were
constantly at war, and their quarrels betrayed symptoms of great
persistence; and if we remember how the observed behaviour of the birds
suggests the fact that they were striving to attain something definite,
we shall understand the nature and extent of the change, and shall, I
fancy, be in a better position to estimate its biological worth at its
true value.
Public-domain text, read in full here on John Shaqi.
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