believe, that we shall find the true explanation of the hostility; it
roughly insures that the number of pairs in any given area does not
exceed the available means of support, and indeed it is difficult to
imagine how such uniformity of distribution as would free the young from
the risk of exposure could be obtained without some such control.
Some birds, however, have no difficulty in finding the necessary food
for their young, yet have great difficulty in finding a station where
they can rear their young in safety; and the area each one occupies has
been reduced to the smallest proportions in order that the maximum
number can be accommodated. Here, any increase in the size of the
territory would inevitably lead to the extinction of the race, so that
nothing stands between failure and success except the ability of the
bird to defend its territory. If we study the bird population at one of
the breeding stations on the coast, we find, generally speaking, that
each kind of bird inhabits a particular portion of the cliff; on the
lower ledges are the Guillemots and Kittiwake Gulls; higher up are
Razorbills and Fulmars, and at the top, where the cliff is broken and
the face of the rock covered with turf and soil, the Puffin finds
shelter for its egg. At the same time there is much overlapping; the
kind of ledge that suits a Razorbill is equally suitable for a Guillemot
or a Fulmar, and so, no matter how successful the Razorbill may be in
establishing a territory and preventing intrusion upon it by other
Razorbills, it will be all to no purpose if it allows itself to be
jostled out of its position by a Fulmar. Hence, inasmuch as breeding
stations are limited and competition for territory so severe, only those
forms in which the fighting instinct responds freely to a wide range of
stimuli will be in a position to maintain a footing upon the cliff.
Public-domain text, read in full here on John Shaqi.
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