Now it is that part of the behaviour routine which has reference to the
relationship between one bird and another upon which, for the time
being, I wish to dwell; for the interest that A displays in B is by no
means always the same--it changes according to the season, and this
change can be observed to be uniform throughout a wide range of species.
In winter, in whatsoever direction we turn, we observe not only that
different individuals but that different species also collect together
in flocks. And since food at that season is not always easy to obtain,
and, moreover, is only to be found in certain situations, which are
limited both in number and extent, it would seem that such assemblages
are in the main determined by accident. No doubt the abundance or the
scarcity of food does determine the movements of birds, and hence to
that extent may be held to account for the flocks. But we shall but
deceive ourselves if we think that it is the sole or even the principal
reason, or that the situation is in no wise affected by internal
factors. The behaviour of the individual in relation to the flock bears
ample testimony to the presence of a gregarious impulse which derives
satisfaction from the fact of close association.
As an illustration, let us take a bird whose movements are easily
watched, and in whose hereditary constitution the impulse to which I
allude seems to be strongly implanted--the Curlew. When the breeding
season is over, Curlew leave the mountain and the moor and return to
the coast or tidal estuaries for the remainder of the year. Here, at low
water, they find an abundant supply of food--crustaceans amongst the
sea-weed upon the rocks, and lobworms (_Arenicola piscatorum_) in the
mud as the tide advances or recedes. But when the tide is full, they
retire to those parts of the shore that remain uncovered--to isolated
rocks, or to sand-dunes, or it may even be to pasture-land in the
neighbourhood. During this period of repose large numbers of individuals
gather together on a comparatively small space of ground. They are not
constrained to do so by any shortage of accommodation, nor by any
question relative to food, nor, for the matter of that, by any
circumstance in the external environment; they are brought together
solely, this at least is the impression that one gains, by some
inherited impulse working towards that end. And their subsequent course
of behaviour tends to confirm that impression. For if we watch the
gathering together of the different units of which the flock is
composed, and study more particularly the emotional manifestation which
accompanies their arrival and departure, we shall find that the coming
of a companion arouses some emotion which is expressed by a vocal
outburst that sweeps through the flock.
Public-domain text, read in full here on John Shaqi.
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