The inference, therefore, is that atmospheric
changes bear some relation to the functioning of the instinct; but
whether it be temperature, or humidity, or the direction and velocity of
the wind, or a combination of two or more of these factors that supplies
the stimulus, we cannot tell.
The appropriate organic condition and the stimulus have then still to be
determined, and we must pursue our inquiry from the point at which the
impulse comes into functional activity. We will take a simple case, and
one free from complication.
Let us suppose that there is an area bereft of bird life, if it can be
so imagined, but in proximity to other inhabited areas. Into this area,
whilst in search of isolation, let us imagine that a Yellow Bunting
finds its way. After the manner of its race it establishes a territory
and occupies, let us say at a low computation, half an acre of ground.
It then obtains a mate, breeds, and rears offspring, two of which we
will assume are males. Reproduction ended, the birds desert the area,
and in the following spring, when the impulse again asserts itself,
parents and offspring seek again their former haunts. We now have three
males, each of which occupies half an acre, and each of which rears two
offspring--that is the position at the close of the second year. In the
third year the number will have increased to nine and the area occupied
to 4½ acres; and so on in succeeding years, until by the beginning of
the eleventh year, we have 59,048 Yellow Buntings occupying 29,524½
acres or 46 square miles. This, then, will be the result of the
operation of the impulse, providing that all the individuals survive and
that no complications supervene.
But of course complications are numerous, some of which retard while
others accelerate the rate of expansion. These complications arise from
various sources--in the first place from natural enemies which prey upon
the birds or upon their eggs; in the next place from climate which, if
it happens to be unfavourable, may mean that food is scarce and that
only a small percentage of the young survive; and lastly from
rivals--and by rivals I mean closely related forms that require a
similar station and similar food--which, by occupying available ground,
may check expansion, or, by forcing a continuation of the search, may
widen it.
Public-domain text, read in full here on John Shaqi.
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