Texas Fossils: An Amateur Collector's HandbookMatthews, William Henry
Science
Texas Fossils: An Amateur Collector's Handbook
Matthews, William Henry
Paleontology -- Texas
Most zoologists and paleontologists recognize three classes of
coelenterates: (1) the Hydrozoa, containing the small animals known as
hydroids, (2) the Scyphozoa, which includes the jellyfish, and (3) the
Anthozoa, which includes the corals and sea anemones. Because of their
extreme fragility and lack of hard parts, hydrozoans and scyphozoans are
not commonly found as fossils. They do, however, have a long geologic
history and may be preserved when unusual conditions of fossilization
occur. The anthozoans, especially the corals, are by far the most
important class geologically, and these forms have left a very good
paleontological record.
CLASS ANTHOZOA.—
This class is composed of a group of exclusively marine organisms and
includes the corals and sea anemones. The coral animal, or _polyp_,
secretes a cup-shaped calcareous (limy) exoskeleton. This skeleton,
called a _corallite_, is usually divided by radial partitions called
_septa_. The polyp lives in the _calyx_, which is the central
bowl-shaped depression in the top of the corallite (fig. 11a).
_Solitary_ corals form an individual corallite for each polyp, and
because of their shape these may be given such names as “horn corals”
(_Lophophyllidium_, Pl. 15) or “button corals” (_Micrabacia_, Pl. 16).
_Colonial_ or _compound_ corals (Pl. 15) live together in colonies,
which are formed of many individual skeletons attached to each other
(fig. 11b), and the compound mass of coral skeletons formed in this
manner is called a _corallum_. Fossil corals commonly occur in many
marine limestones and in places constitute a large portion of the rock.
[Illustration: Fig. 11. Morphology and principal parts of corals. (a)
Solitary or “horn” coral. (b) Colonial or compound coral.]
a
Columella
Septum
Corallite
b
Calyx
Septum
Corallum
The class Anthozoa has been divided into several subclasses, but only
one, the Zoantharia, is of paleontological importance.
Subclass Zoantharia.—
Most corals and all sea anemones belong to this subclass. Zoantharians
are either colonial or solitary and, because most of them possess a hard
preservable exoskeleton, they are the most important group of anthozoans
geologically. The various orders of the subclass Zoantharia are
discussed below.
Order Rugosa.—
These are corals in which the septa are arranged in cycles of four. Both
solitary and colonial forms occur, and they are found only in rocks of
Paleozoic age. Rugose corals are abundant in many of the Paleozoic
formations of Texas, and two of the more typical forms
(_Lophophyllidium_ and _Caninia_) are illustrated in Plate 15. Members
of this order have been placed in the subclass Tetracoralla of older
classifications.
Order Scleractinia.—
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