The secretion came to an end in both stomachs at the same time.
The above is represented in curves in Figs. 2 and 3, the scale on which
that for the main stomach is drawn being ten times less than that for
the small. As you see, the progress of secretion is identical in both.
The existence of a fistula leading into the large stomach affords us
also the possibility of performing an experiment upon our dog which
is exactly the converse of the sham feeding experiment, and which
constitutes a real _experimentum crucis_. While in sham feeding, we
had only, so to speak, the beginning of digestion before us, we are
now able in our cross experiment to start at the continuation of this
beginning. For this purpose it is only necessary to bring the food
into the stomach through the fistula, without attracting the dog’s
attention. Since in this experiment it is above all necessary not to
excite the dog’s appetite, it is best to carry out the procedure on the
sleeping animal. I may add at once, however, that the same result can
be obtained on the waking animal, only the process must be performed
unnoticed, and the animal’s attention must be diverted from thoughts of
food.
[Illustration: FIGURE 2.--Curve of secretion from the miniature
stomach.]
[Illustration:
FIGURE 3.--The same from the main stomach reduced ten times.]
The results of this experiment are striking, and do not in any way
resemble the secretion after normal feeding. Some kinds of food, for
instance bread and coagulated white of the hen’s egg, when directly
introduced into the stomach, do not yield a single drop of juice
during the first hour or more afterwards. This holds good both for
the small and large stomachs. When a glass rod is introduced into the
food contained in the organ it remains dry. Flesh, if introduced at
this stage, is able to excite a secretion, but the appearance of the
juice is considerably retarded. It begins from fifteen to forty-five
minutes after the feeding, instead of from six to ten, is under normal
circumstances extremely scanty during the first hour (3 c.c. to 5 c.c.
instead of 12 c.c. to 15 c.c.), and possesses a very low digestive
power.
Here is an experiment by Dr. Lobassoff:
400 grms. of flesh were brought into the stomach.
Hour. Quantity of juice. Digestive power.
1st 3.7 c.c. 2.0 mm.
2nd 10.6 ” 1.63 ”
3rd 9.2 ” 1.5 ”
4th 7.0 ” 1.88 ”
5th 5.6 ” 2.25 ”
6th 6.6 ” 2.63 ”
7th 7.5 ” 1.88 ”
8th 5.3 ” 2.0 ”
9th 3.0 ” 5.0 ”
10th 0.2 ” — ”
The secretion began twenty-five minutes after introducing the food. I
now ask you to compare the following tables:
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