was seen to open only seven times in twenty minutes following the
arrival of a hard particle of food at the pylorus. The conclusion may
therefore be drawn that hard morsels keep the pylorus closed and hinder
the passage of the food into the duodenum.
3. _Activity of the cardiac portion._--The part played by the fundus
apparently has not hitherto been properly appreciated. It has been
regarded as the place for peptic digestion, or as a passive reservoir
for food; but it is in fact a most interestingly active reservoir.
[Illustration: FIGURE 2.]
[Illustration: FIGURE 3.]
[Illustration: FIGURE 4.]
[Illustration:
FIGURE 5.--Figures 2, 3, 4, and 5 present outlines of the shadow
of the contents of the stomach cast on a fluorescent screen by the
Röntgen rays. The drawings were made by tracing the outline of the
shadow on tissue paper laid upon the fluorescent surface, and are
about one-half the actual size. They show the change in the appearance
of the stomach at intervals for half an hour, from the time of eating
until the stomach is nearly empty.
]
The action of the cardiac portion will be best understood by comparing
the appearances the stomach presents at various stages in a digestive
period. In order to show these stages I carefully made a set of three
tracings of the outlines of the stomach as soon as possible after a
cat had finished eating, and another set of three every half hour
thereafter, until the contents had disappeared (Figs. 2, 3, 4, and
5). These tracings were made by placing white tissue paper over the
fluorescent screen, and drawing with a thick lead pencil, easily seen,
as much of the boundary of the stomach as I could at the end of each
expiration. Between the times for making the drawings the cat was
allowed to rest quietly on a mat, but care was taken to lay her in the
same position on the holder for every drawing. The drawings of each set
were afterwards fastened over one another, so that the lines coincided
as closely as possible. Another piece of tissue paper was then put over
these, and all four sheets were laid on an illuminated pane of glass.
It was thus easy to get a composite tracing which, considering the
movement imparted to the stomach by respiration, and the dimness of
the shadows in the later stages of digestion, probably represents more
exactly than any single drawing the outline of the stomach for each
successive period.
A comparison of these drawings shows that as digestion proceeds the
antrum appears gradually to elongate and acquire a greater capacity,
and that the constrictions make deeper indentations in it. But when the
fundus has lost most of its contents, the longitudinal and circular
fibres of the antrum contract to make it again shorter and smaller.
Its change of form, however, compared with the rest of the stomach, is
slight.
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