The alligator and its alliesReese, A. M. (Albert Moore)
Science
The alligator and its allies
Reese, A. M. (Albert Moore)
Alligators
_The Stomach._ The stomach was sectioned in three regions, as shown
in Figure 35: (1) in the cardiac region very near the opening of the
œsophagus; (2) in the middle or fundic region; and (3) in the region
near the opening of the pylorus. The first two sections are in the
first or large region of the stomach; the third section is in the
second or small region of the stomach (Fig. 35).
The wall as a whole is thickest in the fundus, being there practically
twice as thick as in the pyloric and half again as thick as in the
cardiac region. This great thickening is due mainly to a thickening
of the middle or oblique layer of muscle, which is here remarkably
developed. The mucosa is of nearly uniform thickness in the different
regions and will be described later.
Since there is no striking difference beside that of thickness in the
general structure of the wall of the different regions, the pyloric
region, as seen under low magnification, will now be described (Fig.
45).
The mucosa, _m_, consists of fairly long glands underlaid by a
well-marked muscularis mucosa, _mm_, the latter exhibiting a compact
circular layer over a wider but more scattered layer of longitudinal
fibers. A considerable amount of fibrous connective tissue lies among
the muscle fibers. The circular layer of the muscularis mucosa sends
towards the surface numerous strands or septa between the glands;
six or eight of these are seen in the figure. These strands are not
nearly so numerous in the large region of the stomach. As was said, the
outer or longitudinal layer of the muscularis mucosa is wider but less
compact than the circular and its bundles of fibers are seen in the
figure as a layer of large, scattered dots just beneath the circular
layer.
The submucosa, _sm_, is of average thickness and density. In the fundic
and cardiac regions it seems to extend between the circular and oblique
layers; at any rate, there is a considerable layer of connective tissue
between these two muscular layers.
The circular muscular layer, _cm_, is of only moderate thickness and
is of rather a loose character. In the pyloric region it is not very
distinct from the underlying oblique layer, but in the other regions,
as has just been said, it is separated from the oblique layer by a
considerable layer of connective tissue like that of the submucosa.
The oblique layer, _om_, even in this section of the pyloric region
is the thickest of the three muscle layers; while in the cardiac, and
especially in the fundic, regions it is of great thickness, as was
noted above, and is made up of larger bundles with less intervening
connective tissue.
[Illustration:
FIG. 45. A transsection through the wall of the pyloric region of the
stomach of the feeding animal, under low magnification; _m_, mucosa;
_om_, oblique muscles; other letters as in Figure 42.]
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