The Anatomy of the Human Peritoneum and Abdominal Cavity: Considered from the Standpoint of Development and Comparative AnatomyHuntington, George S. (George Sumner)
Science
The Anatomy of the Human Peritoneum and Abdominal Cavity: Considered from the Standpoint of Development and Comparative Anatomy
Huntington, George S. (George Sumner)
Abdomen; Peritoneum
In the purely carnivorous Marsupials, comprising the family of the
_Dasyuridae_, the reduction of the caecal apparatus, foreshadowed by the
appearance of the distal rudimentary segment as a vermiform appendix in
the wombats, has been carried to the complete elimination of the pouch.
The ileo-colic junction in these animals is simple, marked externally by
a circular constriction and internally by an annular valve. The colon
forms a very short terminal segment of the alimentary canal. The parts
are shown in Fig. 355 in a typical representative of the family,
_Dasyurus viverinus_, the Tasmanian devil.
[Illustration: FIG. 355.--_Dasyurus viverinus_, dasyurus, Tasmanian
devil. Intestinal canal. Ileo-colic junction. (Columbia University
Museum, No. 1463.)]
The structural modifications encountered in the digestive tract of these
carnivorous Marsupials can properly be regarded as the result of their
habitual diet, and we will meet with analogous and identical examples of
caecal reduction in the typical Carnivores among the placental mammals
(cf. p. 212).
III. Subclass: Monodelphia.
III. Order: Edentata.
In all probability the Sloths, Ant-eaters and Armadillos composing this
order represent a highly specialized remnant of an ancient group now
largely extinct. In respect to the ileo-colic junction the Edentates may
be arranged in two groups which offer, within the limited number of
existing species, a very complete transitional series.
I. SYMMETRICAL TYPE OF ILEO-COLIC JUNCTION.
1. _Differentiation in caliber, with direct funnel-like transition of
small into large intestine. No caecum._
Beyond the ileo-colic junction the caliber of the large intestine
increases gradually. The terminal ileum is thus implanted into the apex
of a funnel formed by the proximal segment of the colon.
Examples of this type are furnished by _Myrmecophaga jubata_, the great
ant-eater (Fig. 356), and by _Choloepus didactylus_, the two-toed sloth
(Fig. 357).
[Illustration: FIG. 356.--_Myrmecophaga jubata_, great ant-eater.
Ileo-colic junction. (Columbia University Museum, No. 1519.)]
[Illustration: FIG. 357.--_Choloepus didactylus_, two-toed sloth.
Ileo-colic junction. (Columbia University Museum, No. 714.)]
2. _Abrupt demarcation of small and large intestine, with median
transition of ileum._
The caliber of the intestine enlarges rapidly immediately beyond the
ileo-colic junction. This form is derived from the preceding by the
substitution of the abrupt ileo-colic transition for the gradual
funnel-shaped development of the large intestine.
The type is illustrated by _Tatusia novemcincta_, the nine-banded
armadillo (Fig. 358), and is also found in two other armadillos,
_Tolypeutes_ and _Xenurus_.
[Illustration: FIG. 358.--_Tatusia novemcincta_, nine-banded armadillo.
Ileo-colic junction. (Columbia University Museum, No. 176.)]
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