The Ancient Life History of the Earth: A Comprehensive Outline of the Principles and Leading Facts of Palæontological ScienceNicholson, Henry Alleyne
Science
The Ancient Life History of the Earth: A Comprehensive Outline of the Principles and Leading Facts of Palæontological Science
Nicholson, Henry Alleyne
Paleontology
In another great group of Devonian Ganoids, we meet with fishes
more or less closely allied to the living _Polypteri_ (fig. 105)
of the Nile and Senegal. In this group (fig. 106) the pectoral
fins consist of a central scaly lobe carrying the fin-rays on
both sides, the scales being sometimes rounded and overlapping
(fig. 106), or more commonly rhomboidal and placed edge to edge
(fig. 105, A). Numerous forms of these "Fringe-finned" Ganoids
occur in the Devonian strata, such as _Holoptychius, Glyotoloemus,
Osteolepis, Phaneropleuron_, &c. To this group is also to be
ascribed the huge _Onychodus_ (fig. 102, d and e), with its
large, rounded, overlapping scales, an inch in diameter, and its
powerful pointed teeth. It is to be remembered, however, that
some of these "Fringe-finned" Ganoids are probably referable
to the small but singular group of the "Mud-fishes" (_Dipnoi_),
represented at the present day by the singular _Lepidosiren_
of South America and Africa, and the _Ceratodus_ of the rivers
of Queensland.
[Illustration: Fig. 105.--A, _Polypterus_, a recent Ganoid
fish; B, _Osteolepis_, a Devonian Ganoid; a a, Pectoral fins,
showing the fin-rays arranged round a central lobe.]
[Illusration: Fig. 106.--_Holoptychius nobilissimus_, restored.
Old Red Sandstone, Scotland. A, Scale of the same.]
Leaving the Ganoid fishes, it still remains to be noticed that
the Devonian deposits have yielded the remains of a number of
fishes more or less closely allied to the existing Sharks, Rays,
and _Chimoeroe_ (the _Elasmobranchii_). The majority of the forms
here alluded to are allied not to the true Sharks and Dog-fishes,
but to the more peaceable "Port Jackson Sharks," with their blunt
teeth, adapted for crushing the shells of Molluscs. The collective
name of "Cestracionts" is applied to these; and we have evidence of
their past existence in the Devonian seas both by their teeth, and
by the defensive spines which were implanted in front of a greater
or less number of the fins. These are bony spines, often variously
grooved, serrated, or ornamented, with hollow bases, implanted
in the integument, and capable of being erected or depressed
at will. Many of these "fin-spines" have been preserved to us
in the fossil condition, and the Devonian rocks have yielded
examples belonging to many genera. As some of the true Sharks
and Dog-fishes, some of the Ganoids, and even some Bony Fishes,
possess similar defences, it is often a matter of some uncertainty
to what group a given spine is to be referred. One of these spines,
belonging to the genus _Machoeracanthus_, from the Devonian rocks
of America, has been figured in a previous illustration (fig.
102, f).
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