The Ancient Life History of the Earth: A Comprehensive Outline of the Principles and Leading Facts of Palæontological ScienceNicholson, Henry Alleyne
Science
The Ancient Life History of the Earth: A Comprehensive Outline of the Principles and Leading Facts of Palæontological Science
Nicholson, Henry Alleyne
Paleontology
A few Star-fishes and Brittle-stars are known to occur in the
Carboniferous rocks; but the only other Echinodemls of this period
which need be noticed are the Sea-urchins (_Echinoids_). Detached
plates and spines of these are far from rare in the Carboniferous
deposits; but anything like perfect specimens are exceedingly
scarce. The Carboniferous Sea-urchins agree with those of the
present day in having the body enclosed in a shell formed by
an enormous number of calcareous plates articulated together.
The shell may be regarded as, typically, nearly spherical in
shape, with the mouth in the centre of the base, and the excretory
opening or vent at its summit. In both the ancient forms and the
recent ones, the plates of the shell are arranged in ten zones
which generally radiate from the summit to the centre of the base.
In five of these zones--termed the "ambulacral areas"--the plates
are perforated by minute apertures or "pores," through which
the animal can protrude the little water-tubes ("tube-feet") by
which its locomotion is carried on. In the other five zones--the
so-called "inter-ambulacral areas"--the plates are of larger
size, and are not perforated by any apertures. In all the modern
Sea-urchins each of these ten zones, whether perforate or
imperforate, is composed of two rows of plates; and there are
thus twenty rows of plates in all. In the Palæozoic Sea-urchins,
on the other hand, the "ambulacral areas" are often like those of
recent forms, in consisting of _two_ rows of perforated plates
(fig. 119); but the "inter-ambulacral areas" are always quite
peculiar in consisting each of three, four, five, or more rows
of large imperforate plates, whilst there are sometimes four
or ten rows of plates in the "ambulacral areas" also: so that
there are many more than twenty rows of plates in the entire
shell. Some of the Palæozoic Sea-urchins, also, exhibit a very
peculiar singularity of structure which is only known to exist
in a very few recently-discovered modern forms (viz., _Calveria_
and _Phormosoma_). The plates of the inter-ambulacral areas,
namely, overlap one another in an imbricating manner, so as to
communicate a certain amount of flexibility to the shell; whereas
in the ordinary living forms these plates are firmly articulated
together by their edges, and the shell forms a rigid immovable
box. The Carboniferous Sea-urchins which exhibit this extraordinary
peculiarity belong to the genera _Lepidechinus_ and _Lepidesthes_,
and it seems tolerably certain that a similar flexibility of
the shell existed to a less degree in the much more abundant
genus _Archoeocidaris_. The Carboniferous Sea-urchins, like the
modern ones, possessed movable spines of greater or less length,
articulated to the exterior of the shell; and these structures
are of very common occurrence in a detached condition. The most
abundant genera are _Archoeocidaris_ and _Paloechinus_; but the
characteristic American forms belong principally to _Melonites,
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